Taxonomic Group	Name	Year	Invasion Status	Population Status	Vectors
Cnidarians-Anthozoans	Aiptasiogeton eruptaurantia*	2010	Non-native	Established	Hull Fouling
Fishes	Ambloplites rupestris*	1842	Non-native	Established	None
Fishes	Ameiurus catus	1949	Non-native	Established	None
Fishes	Amia calva*	1942	Non-native	Established	None
Protozoans	Ammonia confertitesta*	2022	Non-native	Established	Ballast Water
Crustaceans-Barnacles	Amphibalanus amphitrite	1967	Non-native	Established	Ballast Water, Hull Fouling, Natural Dispersal
Nematodes	Anguillicola crassus*	1997	Non-native	Established	Ballast Water, Fisheries Accidental (not Oyster)
Arthropoda-Insects	Anisolabis maritima	1901	Non-native	Established	Cargo, Dry Ballast
Algae	Antithamnion hubbsii*	1985	Non-native	Established	Ballast Water, Hull Fouling
Fishes	Aplodinotus grunniens*	1978	Non-native	Established	None
Crustaceans-Copepods	Argulus japonicus	1933	Non-native	Established	None
Tunicates	Ascidiella aspersa	1983	Non-native	Established	Hull Fouling
Entoprocts	Barentsia benedeni	1977	Non-native	Established	Hull Fouling
Algae	Batophora oerstedii*	2015	Non-native	Established	Ballast Water, Fisheries Accidental (not Oyster), Hull Fouling
Mollusks-Gastropods	Bithynia tentaculata	1879	Non-native	Established	None
Cnidarians-Hydrozoans	Blackfordia virginica	1904	Non-native	Established	Ballast Water, Hull Fouling
Annelids-Polychaetes	Boccardiella ligerica	1990	Non-native	Established	Ballast Water
Algae	Bonnemaisonia hamifera	1927	Non-native	Established	Ballast Water, Hull Fouling
Tunicates	Botrylloides violaceus	1980	Non-native	Established	Hull Fouling
Arthropoda-Insects	Brachydeutera longipes*	1983	Non-native	Established	None
Annelids-Oligochaetes	Branchiura sowerbyi*	1955	Non-native	Established	None
Algae	Bryopsis maxima*	2013	Non-native	Established	Ballast Water, Hull Fouling
Bryozoans	Bugula neritina	1985	Non-native	Established	Hull Fouling
Bryozoans	Bugulina simplex	1872	Non-native	Established	Hull Fouling
Annelids-Oligochaetes	Cambarincola pamelae*	2003	Non-native	Established	None
Crustaceans-Amphipods	Caprella mutica	1998	Non-native	Established	Ballast Water, Hull Fouling
Fishes	Carassius auratus	1839	Non-native	Established	None
Crustaceans-Crabs	Carcinus maenas	1817	Non-native	Established	Dry Ballast, Hull Fouling
Algae	Ceramium secundatum*	2007	Non-native	Established	Ballast Water, Hull Fouling
Arthropoda-Insects	Chaetococcus phragmitis*	2000	Non-native	Established	None
Fishes	Channa argus	2003	Non-native	Established	None
Crustaceans-Amphipods	Chelura terebrans	1875	Non-native	Established	Hull Fouling
Arthropoda-Insects	Chilacis typhae*	1986	Non-native	Established	None
Crustaceans-Barnacles	Chthamalus fragilis	1898	Non-native	Established	Ballast Water, Hull Fouling, Oyster Accidental
Tunicates	Ciona intestinalis	1850	Non-native	Established	Hull Fouling
Mollusks-Gastropods	Cipangopaludina chinensis	1925	Non-native	Established	None
Tunicates	Clavelina lepadiformis	2009	Non-native	Established	Hull Fouling
Algae	Codium fragile ssp. Fragile	1957	Non-native	Established	Ballast Water, Hull Fouling, Oyster Accidental
Algae	Colpomenia peregrina*	2012	Non-native	Established	Ballast Water, Hull Fouling
Platyhelminthes	Convoluta convoluta*	1995	Non-native	Established	None
Mollusks-Bivalves	Corbicula fluminea	1971	Non-native	Established	None
Cnidarians-Hydrozoans	Cordylophora caspia	1865	Non-native	Established	Hull Fouling
Crustaceans-Amphipods	Corophium volutator	2001	Non-native	Established	None
Cnidarians-Hydrozoans	Craspedacusta sowerbii	1884	Non-native	Established	None
Platyhelminthes	Cryptocotyle lingua*	1911	Non-native	Established	Dry Ballast, Fisheries Intentional, Hull Fouling, Natural Dispersal
Mollusks-Gastropods	Cuthona perca	1994	Non-native	Established	Ballast Water, Hull Fouling
Algae	Cylindrospermopsis raciborskii*	1977	Non-native	Established	None
Fishes	Cyprinus carpio	1832	Non-native	Established	None
Mollusks-Bivalves	Cyrenoida floridana	1952	Non-native	Established	Canal, Dry Ballast, Unknown Vector
Crustaceans-Cladocerans	Daphnia lumholtzi	1999	Non-native	Established	None
Algae	Dasysiphonia japonica	2009	Non-native	Established	Ballast Water, Hull Fouling
Cnidarians-Anthozoans	Diadumene lineata	1892	Non-native	Established	Hull Fouling, Oyster Accidental
Tunicates	Didemnum vexillum	2000	Non-native	Established	Hull Fouling
Tunicates	Diplosoma listerianum	1975	Non-native	Established	Hull Fouling
Mollusks-Nudibranchs	Doris pseudoargus	2017	Non-native	Established	Ballast Water, Hull Fouling
Fishes	Dorosoma petenense	1953	Non-native	Established	Fisheries Intentional
Mollusks-Bivalves	Dreissena bugensis	2008	Non-native	Established	None
Mollusks-Bivalves	Dreissena polymorpha	1991	Non-native	Established	None
Algae	Dumontia contorta*	1927	Non-native	Established	Ballast Water, Hull Fouling
Tunicates	Ecteinascidia turbinata	1961	Non-native	Established	Hull Fouling
Mollusks-Gastropods	Elimia livescens	1872	Non-native	Established	None
Crustaceans-Crabs	Eriocheir sinensis	2007	Non-native	Established	Ballast Water, Fisheries Accidental (not Oyster), Fisheries Intentional
Fishes	Esox lucius*	1888	Non-native	Established	None
Fishes	Etheostoma blennioides*	2011	Non-native	Established	None
Fishes	Etheostoma zonale*	1984	Non-native	Established	None
Crustaceans-Crayfish	Faxonius rusticus	1996	Non-native	Established	None
Crustaceans-Crayfish	Faxonius virilis	1960	Non-native	Established	None
Annelids-Polychaetes	Ficopomatus enigmaticus	1976	Non-native	Established	Ballast Water, Hull Fouling
Arthropoda-Insects	Galerucella calmariensis*	1992	Non-native	Established	None
Arthropoda-Insects	Galerucella pusilla*	1992	Non-native	Established	None
Fishes	Gambusia affinis	1986	Non-native	Established	None
Cnidarians-Hydrozoans	Gonionemus vertens	1894	Non-native	Established	Ballast Water, Hull Fouling, Oyster Accidental
Algae	Gracilaria vermiculophylla	1998	Non-native	Established	Ballast Water, Hull Fouling, Oyster Accidental
Crustaceans-Amphipods	Grandidierella japonica	2013	Non-native	Established	Ballast Water, Hull Fouling
Algae	Grateloupia turuturu	1996	Non-native	Established	Ballast Water, Hull Fouling
Platyhelminthes	Gyrodactylus anguillae*	1999	Non-native	Established	Ballast Water, Fisheries Accidental (not Oyster)
Protozoans	Haplosporidium nelsoni*	1957	Non-native	Established	Ballast Water, Hull Fouling, Oyster Accidental
Annelids-Polychaetes	Hediste diversicolor	1963	Non-native	Established	None
Crustaceans-Crabs	Hemigrapsus sanguineus	1988	Non-native	Established	Ballast Water
Mollusks-Gastropods	Heterogen japonica	2000	Non-native	Established	None
Bryozoans	Hippoporina indica	2001	Non-native	Established	Hull Fouling
Arthropoda-Insects	Holcocranum saturejae*	1955	Non-native	Established	None
Arthropoda-Insects	Holocranum saturejae*	1955	Non-native	Established	None
Crustaceans-Amphipods	Hourstonius sp.	1994	Non-native	Established	Ballast Water, Hull Fouling
Arthropoda-Insects	Hylobius transversovittatus*	1991	Non-native	Established	None
Crustaceans-Isopods	Ianiropsis serricaudis	1999	Non-native	Established	Ballast Water, Hull Fouling
Fishes	Ichthyomyzon unicuspis*	1974	Non-native	Established	None
Fishes	Ictalurus furcatus	1974	Non-native	Established	None
Fishes	Ictalurus punctatus	1889	Non-native	Established	None
Mollusks-Bivalves	Ischadium recurvum	1869	Non-native	Established	Oyster Accidental
Annelids-Polychaetes	Janua heterostropha	2000	Non-native	Established	Hull Fouling
Fishes	Labidesthes sicculus*	1991	Non-native	Established	None
Arthropoda-Insects	Lasioptera hungarica*	2000	Non-native	Established	None
Fishes	Lepomis cyanellus	1900	Non-native	Established	None
Fishes	Lepomis gulosus	1897	Non-native	Established	None
Fishes	Lepomis macrochirus	1855	Non-native	Established	None
Fishes	Lepomis megalotis*	1923	Non-native	Established	None
Fishes	Lepomis microlophus	1974	Non-native	Established	None
Crustaceans-Isopods	Ligia exotica	1924	Non-native	Established	Cargo, Dry Ballast
Arthropoda-Insects	Lipara pullitarsis*	2002	Non-native	Established	None
Arthropoda-Insects	Lipara rufitarsis*	2000	Non-native	Established	None
Arthropoda-Insects	Lipara similis*	1998	Non-native	Established	None
Mollusks-Gastropods	Littorina littorea	1875	Non-native	Established	Dry Ballast, Hull Fouling, Natural Dispersal
Algae	Lomentaria clavellosa*	1972	Non-native	Established	Ballast Water, Hull Fouling
Algae	Lomentaria orcadensis*	1853	Non-native	Established	Hull Fouling
Bryozoans	Lophopodella carteri	1972	Non-native	Established	Aquatic Plant Shipments, Fisheries Accidental (not Oyster), Natural Dispersal
Entoprocts	Loxosomatoides laevis	1995	Non-native	Established	Hull Fouling
Crustaceans-Barnacles	Loxothylacus panopaei	1964	Non-native	Established	Oyster Accidental
Crustaceans-Shrimp	Lysmata vittata	2013	Non-native	Established	Ballast Water
Cnidarians-Hydrozoans	Maeotias marginata	1968	Non-native	Established	Ballast Water, Hull Fouling
Algae	Melanosiphon intestinalis*	2002	Non-native	Established	Ballast Water, Hull Fouling
Algae	Melanothamnus japonicus*	2007	Non-native	Established	Ballast Water, Hull Fouling
Bryozoans	Membranipora membranacea	1990	Non-native	Established	Ballast Water, Hull Fouling, Natural Dispersal
Crustaceans-Amphipods	Microdeutopus gryllotalpa	1872	Non-native	Established	Hull Fouling
Fishes	Micropterus dolomieu	1855	Non-native	Established	None
Fishes	Micropterus punctulatus	2005	Non-native	Established	None
Fishes	Micropterus salmoides	1850	Non-native	Established	None
Fishes	Misgurnus anguillicaudatus*	2013	Non-native	Established	None
Cnidarians-Hydrozoans	Moerisia lyonsi	1965	Non-native	Established	Ballast Water, Hull Fouling
Fishes	Morone chrysops*	1976	Non-native	Established	None
Fishes	Moxostoma macrolepidotum*	1999	Non-native	Established	None
Mollusks-Gastropods	Myosotella myosotis	1832	Non-native	Established	Cargo, Dry Ballast
Mollusks-Bivalves	Mytilopsis leucophaeata	1937	Non-native	Established	Ballast Water, Hull Fouling
Arthropoda-Insects	Nacerdes melanura*	1885	Non-native	Established	Cargo, Hull Fouling
Annelids-Polychaetes	Neodexiospira brasiliensis	1982	Non-native	Established	Hull Fouling
Fishes	Neogobius melanostomus*	2021	Non-native	Established	None
Algae	Neopyropia katadae*	2005	Non-native	Established	Ballast Water, Fisheries Accidental (not Oyster), Hull Fouling
Algae	Neopyropia koreana*	2003	Non-native	Established	Ballast Water, Hull Fouling
Algae	Neopyropia yezoensis*	2005	Non-native	Established	Ballast Water, Fisheries Accidental (not Oyster), Hull Fouling
Fishes	Notropis atherinoides*	1936	Non-native	Established	None
Mollusks-Bivalves	Ostrea edulis	1949	Non-native	Established	Ballast Water, Natural Dispersal, Oyster Intentional
Crustaceans-Shrimp	Palaemon elegans	2017	Non-native	Established	Ballast Water, Discarded Bait, Natural Dispersal
Crustaceans-Shrimp	Palaemon macrodactylus	2001	Non-native	Established	Ballast Water
Fishes	Percina peltata*	1936	Non-native	Established	None
Protozoans	Perkinsus marinus*	1949	Non-native	Established	Oyster Accidental
Arthropoda-Insects	Petrobius brevistylis*	1979	Non-native	Established	Cargo, Dry Ballast
Algae	Phycocalidia suborbiculata*	1966	Non-native	Established	Ballast Water, Hull Fouling
Fishes	Pimephales promelas	1936	Non-native	Established	None
Mollusks-Bivalves	Pisidium amnicum	1962	Non-native	Established	Ballast Water, Canal, Dry Ballast, Packing Material
Mollusks-Gastropods	Pleurocera canaliculata	1869	Non-native	Established	None
Fishes	Pomoxis annularis	1894	Non-native	Established	None
Fishes	Pomoxis nigromaculatus	1873	Non-native	Established	None
Mollusks-Gastropods	Potamopyrgus antipodarum	2021	Non-native	Established	None
Annelids-Oligochaetes	Potamothrix vejdovskyi*	1974	Non-native	Established	None
Crustaceans-Mysids	Praunus flexuosus	2010	Non-native	Established	Ballast Water, Natural Dispersal
Crustaceans-Crayfish	Procambarus clarkii	1981	Non-native	Established	None
Crustaceans-Crayfish	Procambarus zonangulus	2006	Non-native	Established	None
Arthropoda-Insects	Procanace dianneae*	1987	Non-native	Established	Cargo, Hull Fouling
Platyhelminthes	Pseudodactylogyrus anguillae*	1999	Non-native	Established	Ballast Water, Fisheries Accidental (not Oyster)
Fishes	Pylodictis olivaris	1965	Non-native	Established	None
Mollusks-Bivalves	Rangia cuneata	1957	Non-native	Established	Ballast Water, Canal, Oyster Accidental
Mollusks-Gastropods	Rapana venosa	1998	Non-native	Established	Ballast Water, Hull Fouling
Arthropoda-Insects	Rhizedra lutosa*	1988	Non-native	Established	None
Algae	Rhodymenia delicatula*	1996	Non-native	Established	Ballast Water, Hull Fouling
Fishes	Salmo trutta*	1885	Non-native	Established	Fisheries Intentional
Fishes	Sander vitreus*	1879	Non-native	Established	None
Fishes	Scardinius erythrophthalmus*	1986	Non-native	Established	None
Arthropoda-Insects	Sclerocona acutella*	1984	Non-native	Established	None
Crustaceans-Amphipods	Stenothoe gallensis species complex	1960	Non-native	Established	Ballast Water, Hull Fouling
Arthropoda-Insects	Stomoxys calcitrans*	1781	Non-native	Established	None
Mollusks-Gastropods	Stramonita canaliculata	1955	Non-native	Established	Hull Fouling, Oyster Accidental
Algae	Striaria attenuata*	1848	Non-native	Established	Hull Fouling
Tunicates	Styela canopus	1871	Non-native	Established	Hull Fouling
Tunicates	Styela clava	1973	Non-native	Established	Hull Fouling
Tunicates	Styela plicata	2002	Non-native	Established	Hull Fouling
Crustaceans-Isopods	Synidotea laticauda	1999	Non-native	Established	Ballast Water, Hull Fouling
Arthropoda-Insects	Telmatogeton japonicus*	1947	Non-native	Established	Cargo, Hull Fouling
Mollusks-Bivalves	Teredo navalis	1862	Non-native	Established	Hull Fouling
Arthropoda-Insects	Tetramesa phragmitis*	2000	Non-native	Established	None
Arthropoda-Insects	Thambemyia borealis*	2015	Non-native	Established	None
Mollusks-Gastropods	Thecacera pennigera	2003	Non-native	Established	Ballast Water, Hull Fouling
Bryozoans	Tricellaria inopinata	2010	Non-native	Established	Hull Fouling
Algae	Ulonema rhizophorum*	2002	Non-native	Established	None
Algae	Ulva australis	2011	Non-native	Established	None
Algae	Ulva laetevirens*	2011	Non-native	Established	Ballast Water, Hull Fouling
Fishes	Umbra limi*	1974	Non-native	Established	None
Mollusks-Gastropods	Valvata piscinalis	1985	Non-native	Established	None
Bryozoans	Victorella pavida	1920	Non-native	Established	Hull Fouling
Mollusks-Gastropods	Viviparus georgianus	1895	Non-native	Established	None

* = Species summary still under review.

na-et3 (Cape Cod to Cape Hatteras) Summary