Invasion History
First Non-native North American Tidal Record: 1905First Non-native West Coast Tidal Record: 1905
First Non-native East/Gulf Coast Tidal Record: 2005
General Invasion History:
Amathia verticillata is an erect, soft-bodied bryozoan, first described from the Mediterranean Sea (Naples, Italy) and now widespread in tropical, subtropical, and warm-temperate waters around the world (Winston 1977; Gordon and Mawatari 1992) We consider it cryptogenic in the tropical-subtropical Atlantic from North Carolina and Bermuda to Santos, Brazil, (Osburn 1914; Marcus 1937; Osburn 1940; Maturo 1958; Winston 1982, but see Farrapeira, 2011) and the coast of west Africa (Senegal and Gana, Cook 1968), Pacific coast of Central America (Soule et al. 1980, cited by Carlton and Cohen 1995), Arabian Sea and Bay of Bengal waters of India (Robertson 1921, cited by Cohen and Carlton 1995); Karande (1968, cited by Winston 1977; Rao and Ganapati 1978), and southern China, in the South China Sea (Huang 2001).
The origin of this bryozoan is unknown, but it is a documented invader in California (1905, Cohen and Carlton 1995), Hawaii (before 1921, Carlton and Eldredge 2007), New Zealand (Gordon and Mawatari 1992), Australia (before 1975, Keough and Ross 1999), Japan (early 1900s, Asakura 1992); South Korea (Je et al. 1988), the Azores (2008, Amat and Tempera 2009), Madeira (Wirtz and Canning-Clode 2009), and the Canary Islands (2010, Canning-Clode et al. 2013; Minchin 2012), and Taiwan (Minchin et al. 2016). Its larval period is brief (Hyman 1959), so that fouling and ballast water (as fragments of adult colonies or attached to bits of flotsam) are likely vectors for its introduction. A recent review documents the probable introduction of this bryozoan to the Mediterranean Sea, where it was first discovered in the early 19th century but is largely restricted to harbors and marinas (Marchini et al. 2015). Galil and Gevli (2014) argued for a Caribbean origin of A. verticillata, because of its strong association with corals, rocks, and seagrasses there, and the presence of a specialized nudibranch predator there (Okenia zoobotryon).
North American Invasion History:
Invasion History on the West Coast:
Robertson (1905, cited by Cohen and Carlton 1995) observed dense growths of Amathia verticillata in San Diego Bay, where it is still abundant in eelgrass beds (Williams 2007). It is found in Mazatlan, Mexico (Alvarez-Leon and Banta 1984), and the Gulf of California (in 1958, Soule 1963). The pattern of its spread along the California coast is unknown, because most of the records in smaller harbors and bays are quite recent. It was found in Mission Bay, La Jolla, and Newport Bay by 1963 (Soule 1963), and Los Angeles Harbor 'in recent years' (Soule et al. 1980, cited by Cohen and Carlton 1995). In 1993, it appeared in Redwood Creek, South San Francisco Bay (Cohen and Carlton 1995), but has not been seen since (James T. Carlton, personal communication 2013). In a 2001 survey, it was found in small harbors in southern California, including Oceanside Harbor, Channel Islands Harbor, and Avalon Harbor, on Catalina Island (Fairey et al. 2002).
Invasion History on the East Coast:
Amathia verticillata was collected in the Dry Tortugas, Florida, as early as 1908 (Osburn 1914), although it was earlier collected in Bermuda in 1876 (Maturo and Schopf, 1968). It has been treated as native, from North Carolina to Venezuela, by most writers (Osburn 1914; Osburn 1940; Weiss 1948; Maturo 1957; Winston 1982; Pederson and Peterson 2002). However, Carlton and Ruckelshaus (1997) and Gossett et al. (2004) list it as an introduction to Florida and the Gulf of Mexico. We consider it cryptogenic here. A colony found in 2005 at the restoration shipyard of Mystic Seaport, Connecticut, on Long Island Sound (James T. Carlton, personal communication), was an introduction by boat fouling. It was collected in the Mystic River again in 2010 (James T. Carlton, personal communication). It was collected in the Mystic River again in 2010 (James T. Carlton, personal communication). Additional occurrences have been seen from 2014 to 2016, from Wachapreague and Kiptopeake, Virginia and from New Bedford, Massachusetts. Establishment of this warm-water bryozoan in Long Island Sound is unlikely until further warming occurs.
Invasion History in Hawaii:
Robertson (1921) reported having received specimens of A. verticillata from Honolulu Harbor (Robertson 1921, cited by Carlton and Eldredge 2009). It was present in Pearl Harbor by 1921, and in Kaneohe Bay by 1939 (Carlton and Eldredge, 2009), and is also known from Maui (Coles et al. 2004). Recent reports have A. verticillata rapidly becoming abundant at Palmyra Island in the Line Islands, about 2000 km south of Hawaii (Knapp et al. 2011).
Invasion History Elsewhere in the World:
Amathia verticillata is a recent introduction in the Indian Ocean, the South China Sea, and Japan, appearing in the early 1900s, and first collected in Tokyo Bay in 1963 (Asakura 1992; Jebakumar et al. 2017). This bryozoan was newly recorded on the south coast of South Korea, where it became a pest fouling cultured pearl oysters (Je et al. 1988). However, Otani (2006) lists it as a cryptogenic species in Japan. It was first collected in Auckland Harbor, New Zealand about 1960 and has since been found in several harbors on the North Island (Gordon and Mawatari 1992). Amathia verticillata was collected in New South Wales before 1975 (Russ and Wake 1975, cited by Keough and Ross 1999) and in South and Western Australia before 1982 (Bock 1982, cited by Keough and Ross 1999; Huisman et al. 2008).
Amathia verticillata was first reported from Europe in Cadiz, Spain, in 1807, and described from Naples in 1822 Marchini et al. (2015) has described this species as 'pseudo-indigenous', probably introduced very early, but still largely confined to harbors and marinas, and preferring artificial substrates (Marchini et al. 2015). Early Mediterranean records, from west to east include Malaga, Spain, 1823, Algiers, Algeria 1848, Livorno, Italy 1848, Naples, Italy 1822 and Alexandria, Egypt 1828 (Marchini et al. 2015). Locally in the Mediterranean, A. verticillata rapidly colonize marinas, and provide habitat for other introduced species (Marchini et al. 2015). A. verticillata is a recent invader in the Azores, where it appeared in marinas in 2008 on three of the islands (Amat and Tempera 2009), and Madeira, where it was first collected in May of 2009 (Wirtz and Canning-Clode 2009).
In the Eastern Pacific, Amathia verticillata was first collected in Los Frailes Bay in the Gulf of California in 1958,), Baja California and at Mazatlán, Sinaloa (Soule 1963). This bryozoan was collected further south in Oaxaca in 2017 and 2018 (Humara-Gil, et al. 2019). It was first reported from the Pacific coast of South America in 2014, in the Salinas Yacht Club, Ecuador (Priscilla Martinez, personal communication, February 2015, cited by McCann et al. 2015).
Description
Amathia verticillata is an erect, un-calcified bryozoan, with colonies made of irregularly thick transparent, branching or triple-quadruple branching stolons with zooids attached. Colonies can resemble ‘stringy, gelatinous noodles or sauerkraut up to 2 m long’ (Gordon and Mawatari 1992). The zooids are ovoid, about 380 X 580 µm in size, and occur in clusters on either side of the thick transparent stolon. The polypides have eight tentacles and large gizzards. The zooids brood whitish embryos (description from Winston 1982; Gordon and Mawatari 1992; Vieira et al. 2014). Colonies vary greatly in size, 400–700 mm in Brazil (Farrapeira et al. 2010) to 2200 mm (Amat et al. 2011).
Recent molecular work has confirmed that Amathia verticillata is a single global species (Nascimento et al. 2021). A recent genetic analysis of the family Vesiculariidae supports a synonymy of the genera Amathia, Bowerbankia, and Zoobotryon (Waeschenbach et al. 2015).
Taxonomy
Taxonomic Tree
Kingdom: | Animalia | |
Phylum: | Bryozoa | |
Class: | Gymnolaemata | |
Order: | Ctenostomata | |
Suborder: | Carnosa | |
Superfamily: | Vesicularioidea | |
Family: | Vesiculariidae | |
Genus: | Amathia | |
Species: | verticillata |
Synonyms
Hydra verticillata (Della Chaije, 1828)
Zoobotryon pellucidus (Ehrenberg, 1839)
Amathia verticillata (Waeschenbach et al., 2015)
Potentially Misidentified Species
Ecology
General:
Life History- Amathia verticillata is an erect, uncalcified bryozoan, composed of many individual zooids. The zooids feed by extending the ciliated tentacles of the lophophore as a funnel, creating a current, and driving food particles into their mouths. The food is guided along the tentacles and through the pharynx by the cilia. Larger food particles can be moved or captured by flicking or contracting the tentacles. The zooids are hermaphroditic, and produce large yolky eggs, which hatch into lecithotrophic larvae, which are planktonic for short periods (less than 1 day). Larvae settle on a substrate and metamorphose into the first zooid of a colony, an ancestrula (Barnes 1983).
Ecology- This bryozoan is found on wood, rock, vegetation (including eelgrass and mangroves), shells, and man-made structures, including vessel hulls (Winston 1982; Gordon and Mawatari 1992; Minchin 2012). In an investigation of chemical compounds affecting feeding on A. verticillata, a Brazilian population had alkaloids that deterred feeding, while Florida populations had variable resistance to feeding. Dos Santos et al. (2017) suggest that the success of this bryozoan is due to factors other than toxicity.
Food:
Phytoplankton
Trophic Status:
Suspension Feeder
SusFedHabitats
General Habitat | Marinas & Docks | None |
General Habitat | Rocky | None |
General Habitat | Grass Bed | None |
General Habitat | Coarse Woody Debris | None |
General Habitat | Oyster Reef | None |
General Habitat | Canals | None |
General Habitat | Mangroves | None |
General Habitat | Vessel Hull | None |
Salinity Range | Mesohaline | 5-18 PSU |
Salinity Range | Polyhaline | 18-30 PSU |
Salinity Range | Euhaline | 30-40 PSU |
Tidal Range | Subtidal | None |
Vertical Habitat | Epibenthic | None |
Life History
Tolerances and Life History Parameters
Minimum Salinity (‰) | 15 | Nair et al. 1992 |
Maximum Salinity (‰) | 40 | Field salinity (Shark Bay, Western Australia) (Wyatt et al. 2005) |
Minimum Length (mm) | 100 | Colony length, Azores (Amat et al. 2009, on marina floats) |
Maximum Height (mm) | 2,200 | Colony length, Azores (Amat et al. 2009, on marina floats) |
Broad Temperature Range | None | Warm temperate-Tropical |
Broad Salinity Range | None | Mesohaline-Euhaline |
General Impacts
Economic Impacts
Shipping/Boating- Amathia verticillata is known from ship fouling (Woods Hole Oceanographic Institution 1952) and is reported to cause fouling problems to boats and marinas in southern California (Johnson et al. 2006).
Fisheries- Amathia verticillata causes major problems by clogging shrimp-fishing gear in Galveston Bay, Texas (Gossett et al. 2004). It was reported as a pest species fouling cultured pearl oysters, Pinctada fuca, on the south coast of Korea (Je et al. 1988).
Ecological Impacts
Habitat Change- Amathia verticillata grows epiphytically on the leaves of eelgrass (Zostera marina), killing plants in San Diego Bay, which creates gaps in the seagrass canopy, leaving bare patches of sediment available for colonization by other algal species (Williams 2007).
Regional Impacts
NEP-VI | Pt. Conception to Southern Baja California | Economic Impact | Shipping/Boating | ||
Amathia verticillata is reported to cause fouling problems to boats and marinas in southern California (Johnson et al. 2006). | |||||
NEP-VI | Pt. Conception to Southern Baja California | Ecological Impact | Competition | ||
Amathia verticillata grows epiphytically on the leaves of eelgrass (Zostera marina) in San Diego Bay, killing them (Williams 2007). |
|||||
NEP-VI | Pt. Conception to Southern Baja California | Ecological Impact | Habitat Change | ||
Amathia verticillata kills eelgrass plants in San Diego Bay, creating gaps in the seagrass canopy and bare patches of sediment, which are then colonized by adult plants (Williams 2007). | |||||
P020 | San Diego Bay | Ecological Impact | Competition | ||
Amathia verticillata grows epiphytically on the leaves of eelgrass (Zostera marina) in San Diego Bay, killing them (Williams 2007). | |||||
P020 | San Diego Bay | Ecological Impact | Habitat Change | ||
Amathia verticillata kills eelgrass plants in San Diego Bay, creating gaps in the seagrass canopy and bare patches of sediment, which are then colonized by adult plants (Williams 2007). | |||||
CA | California | Ecological Impact | Competition | ||
Amathia verticillata grows epiphytically on the leaves of eelgrass (Zostera marina) in San Diego Bay, killing them (Williams 2007). | |||||
CA | California | Ecological Impact | Habitat Change | ||
Amathia verticillata kills eelgrass plants in San Diego Bay, creating gaps in the seagrass canopy and bare patches of sediment, which are then colonized by adult plants (Williams 2007). |
Regional Distribution Map
Bioregion | Region Name | Year | Invasion Status | Population Status |
---|---|---|---|---|
P062 | _CDA_P062 (Calleguas) | 2001 | Non-native | Established |
P058 | _CDA_P058 (San Pedro Channel Islands) | 2001 | Non-native | Established |
P023 | _CDA_P023 (San Louis Rey-Escondido) | 2001 | Non-native | Established |
P090 | San Francisco Bay | 1993 | Non-native | Established |
NEP-V | Northern California to Mid Channel Islands | 1980 | Non-native | Established |
P050 | San Pedro Bay | 1980 | Non-native | Established |
P030 | Mission Bay | 1963 | Non-native | Established |
P022 | _CDA_P022 (San Diego) | 1963 | Non-native | Established |
P040 | Newport Bay | 1959 | Non-native | Established |
P020 | San Diego Bay | 1905 | Non-native | Established |
NEP-VI | Pt. Conception to Southern Baja California | 1905 | Non-native | Established |
Occurrence Map
OCC_ID | Author | Year | Date | Locality | Status | Latitude | Longitude |
---|---|---|---|---|---|---|---|
697232 | Maloney et al. 2007 | 2005 | 2005-04-27 | Kelco Dock | Non-native | 32.6927 | -117.1470 |
697339 | Introduced Species Study | 2006 | 2006-08-24 | New Site (Blue Awnings) | Non-native | 33.7224 | -118.0561 |
697422 | Introduced Species Study | 2006 | 2006-09-14 | Newport Bay Harbor Entrance | Non-native | 33.5974 | -117.8798 |
697515 | Introduced Species Study | 2006 | 2006-08-25 | Northern Corner of Harbor | Non-native | 33.9830 | -118.4465 |
697659 | Introduced Species Study | 2006 | 2006-09-15 | East of Bridge | Non-native | 33.4591 | -117.6992 |
697740 | Introduced Species Study | 2006 | 2006-08-24 | Yellow Condos | Non-native | 33.7283 | -118.0602 |
697768 | Maloney et al. 2007 | 2005 | 2005-06-29 | San Diego Bay Commercial Fishing Fleet | Non-native | 32.7109 | -117.1739 |
697770 | ISS 2000-2002 Survey Data | 2001 | 2001-10-09 | San Diego Bay Commercial Fishing Fleet | Non-native | 32.7109 | -117.1739 |
697808 | Cohen et al. 2002 | 2000 | 2000-08-24 | Watchorn Basin | Non-native | 33.7203 | -118.2764 |
697860 | Introduced Species Study | 2011 | 2011-05-04 | Ski Islands Marina | Non-native | 32.7939 | -117.2232 |
697867 | Introduced Species Study | 2006 | 2006-09-12 | Ski Islands Marina | Non-native | 32.7939 | -117.2232 |
698112 | Introduced Species Study | 2011 | 2011-05-04 | Hilton Resort Dock | Non-native | 32.7788 | -117.2127 |
698119 | Introduced Species Study | 2006 | 2006-09-12 | Hilton Resort Dock | Non-native | 32.7788 | -117.2127 |
698534 | Maloney et al. 2007 | 2005 | 2005-04-27 | Glorietta Bay | Non-native | 32.6796 | -117.1740 |
698673 | Introduced Species Study | 2011 | 2011-05-03 | Marine Terminal (Paco) | Non-native | 32.6584 | -117.1191 |
698677 | Maloney et al. 2007 | 2005 | 2005-04-27 | Marine Terminal (Paco) | Non-native | 32.6584 | -117.1191 |
698714 | Introduced Species Study | 2006 | 2006-07-25 | Private Dock | Non-native | 34.1798 | -119.2297 |
698780 | Maloney et al. 2007 | 2005 | 2005-06-28 | Navy - 28th St. | Non-native | 32.6836 | -117.1311 |
698840 | Introduced Species Study | 2006 | 2006-09-13 | Guest Dock | Non-native | 33.2091 | -117.3947 |
698967 | Introduced Species Study | 2011 | 2011-05-04 | Dana Inn Marina | Non-native | 32.7671 | -117.2362 |
698970 | Introduced Species Study | 2006 | 2006-09-12 | Dana Inn Marina | Non-native | 32.7671 | -117.2362 |
699150 | Cohen et al. 2002 | 2000 | 2000-08-30 | Rainbow Lagoon | Non-native | 33.7631 | -118.1914 |
699157 | ISS 2000-2002 Survey Data | 2001 | 2001-07-10 | Channel Islands Harbor Epifaunal 02 | Non-native | 34.1811 | -119.2319 |
699182 | ISS 2000-2002 Survey Data | 2001 | 2001-08-14 | Oceanside Epifaunal 01 | Non-native | 33.2118 | -117.3951 |
699274 | Introduced Species Study | 2010 | 2010-06-02 | Oakland Inner Harbor - Shipping cranes | Non-native | 37.7947 | -122.3095 |
699588 | ISS 2000-2002 Survey Data | 2001 | 2001-08-16 | Long Beach Epifaunal 02 | Non-native | 33.7477 | -118.1975 |
699643 | Introduced Species Study | 2010 | 2010-07-12 | Pier 45 | Non-native | 37.8111 | -122.4196 |
699694 | Introduced Species Study | 2006 | 2006-08-25 | Vessel Assist Dock | Non-native | 33.9783 | -118.4569 |
699729 | Introduced Species Study | 2011 | 2011-05-04 | Seaforth | Non-native | 32.7621 | -117.2365 |
699732 | Introduced Species Study | 2006 | 2006-09-12 | Seaforth | Non-native | 32.7621 | -117.2365 |
699806 | Maloney et al. 2007 | 2005 | 2005-06-29 | Coronado Wharf | Non-native | 32.6992 | -117.1684 |
700023 | Introduced Species Study | 2010 | 2010-07-12 | Cruise Ship Pier | Non-native | 37.8085 | -122.4060 |
700110 | Introduced Species Study | 2011 | 2011-05-05 | Middle Harbor Yacht Slip | Non-native | 33.2106 | -117.3960 |
700197 | Introduced Species Study | 2006 | 2006-08-25 | Marina del Rey Harbor Entrance | Non-native | 33.9702 | -118.4496 |
700249 | ISS 2000-2002 Survey Data | 2001 | 2001-07-12 | Huntington Harbour Epifaunal 04 | Non-native | 33.7278 | -118.0787 |
700380 | Introduced Species Study | 2006 | 2006-09-15 | Dana Point Harbor Mouth Slip | Non-native | 33.4594 | -117.6941 |
700401 | Introduced Species Study | 2006 | 2006-08-24 | Near Huntington Launch Ramp | Non-native | 33.7279 | -118.0786 |
700547 | ISS 2000-2002 Survey Data | 2001 | 2001-07-10 | Channel Islands Harbor Epifaunal 05 | Non-native | 34.1739 | -119.2236 |
700648 | Introduced Species Study | 2006 | 2006-09-12 | Santa Barbara Cove | Non-native | 32.7774 | -117.2484 |
700684 | Introduced Species Study | 2006 | 2006-08-24 | Private Beach | Non-native | 33.7175 | -118.0659 |
700984 | A. Sewell, pers. comm. 1995, in Cohen and Carlton 1995 | 1995 | Mission Bay | Non-native | 32.7791 | -117.2288 | |
701397 | Introduced Species Study | 2006 | 2006-09-13 | Oceanside Commercial Fishing Dock | Non-native | 33.2057 | -117.3897 |
701398 | Introduced Species Study | 2011 | 2011-05-05 | Oceanside Commercial Fishing Dock | Non-native | 33.2057 | -117.3897 |
702049 | ISS 2000-2002 Survey Data | 2001 | 2001-07-11 | Los Angeles Epifaunal 04 | Non-native | 33.7655 | -118.2529 |
702115 | Maloney et al. 2007 | 2005 | 2005-06-28 | Switzer Creek (Dole) | Non-native | 32.7017 | -117.1585 |
702203 | Maloney et al. 2007 | 2005 | 2005-04-27 | Chula Vista Marina | Non-native | 32.6225 | -117.1023 |
702481 | Robertson 1905 | 1905 | San Diego Bay | Non-native | 32.6717 | -117.1439 | |
702482 | A. Sewell, pers. comm. 1995, in Cohen and Carlton 1995 | 1995 | San Diego Bay | Non-native | 32.6717 | -117.1439 | |
702880 | Introduced Species Study | 2006 | 2006-08-25 | NW Corner of Harbor | Non-native | 33.9830 | -118.4564 |
703073 | Introduced Species Study | 2011 | 2011-05-03 | San Diego Bay Cruise Ship Terminal | Non-native | 32.7168 | -117.1759 |
703154 | ISS 2000-2002 Survey Data | 2001 | 2001-07-12 | Huntington Harbour Epifaunal 02 | Non-native | 33.7175 | -118.0658 |
703347 | Maloney et al. 2007 | 2005 | 2005-04-27 | Coronado Cays | Non-native | 32.6274 | -117.1329 |
703451 | Introduced Species Study | 2006 | 2006-09-14 | Back Bay Marina | Non-native | 33.6194 | -117.8933 |
703499 | Introduced Species Study | 2006 | 2006-09-13 | North Corner of Harbor | Non-native | 33.2122 | -117.3954 |
703537 | Introduced Species Study | 2006 | 2006-09-14 | Balboa Coves | Non-native | 33.6213 | -117.9364 |
703760 | Introduced Species Study | 2006 | 2006-09-14 | Balboa North Channel | Non-native | 33.6097 | -117.8957 |
704210 | Introduced Species Study | 2006 | 2006-09-14 | Lido Peninsula | Non-native | 33.6082 | -117.9195 |
704233 | Introduced Species Study | 2006 | 2006-08-24 | Pump-A-Head Dock | Non-native | 33.7026 | -118.0542 |
704351 | Introduced Species Study | 2011 | 2011-04-21 | Super Mexico Pier | Non-native | 33.7708 | -118.2113 |
704379 | Cohen et al. 2002 | 2000 | 2000-08-24 | Newmarks Yacht Harbor | Non-native | 33.7644 | -118.2497 |
704447 | K. Wasson pers. comm., cited in Cohen and Carlton 1995 | 1993 | Redwood Creek | Non-native | 37.5066 | -122.2147 | |
704648 | Maloney et al. 2007 | 2005 | 2005-06-28 | Navy - Carrier Base | Non-native | 32.7062 | -117.1886 |
717023 | Soule 1963 | 1959 | Newport Harbor | Non-native | 33.6092 | -117.9067 | |
717025 | Soule 1963 | 1963 | La Jolla | Non-native | 32.8479 | -117.2776 | |
717026 | Soule 1963 | 1963 | Mission Bay | Non-native | 32.7791 | -117.2288 | |
758036 | Bullivant 1968 | 1965 | Newport Harbor | Non-native | 33.6092 | -117.9067 | |
758037 | de Rivera et al. 2005 | 2003 | Dana Marina, Mission Bay | Non-native | 32.7675 | -117.2365 | |
758038 | de Rivera et al. 2005 | 2003 | Seaforth Marina, Mission Bay | Non-native | 32.7630 | -117.2373 | |
758039 | de Rivera et al. 2005 | 2003 | Chula Vista Marina, San Diego Bay | Non-native | 32.6244 | -117.1038 | |
758040 | de Rivera et al. 2005 | 2003 | Sunroad Marina, San Diego Bay | Non-native | 32.7263 | -117.1922 | |
767418 | Ruiz et al., 2015 | 2013 | 2013-07-19 | SeaWorld Marina, Mission Bay, CA, California, USA | Non-native | 32.7676 | -117.2314 |
767451 | Ruiz et al., 2015 | 2013 | 2013-07-29 | Mission Bay Yacht Club, Mission Bay, CA, California, USA | Non-native | 32.7778 | -117.2485 |
767471 | Ruiz et al., 2015 | 2013 | 2013-08-04 | Bahia Resort Marina, Mission Bay, CA, California, USA | Non-native | 32.7731 | -117.2478 |
767489 | Ruiz et al., 2015 | 2013 | 2013-07-31 | Campland on the Bay, Mission Bay, CA, California, USA | Non-native | 32.7936 | -117.2234 |
767501 | Ruiz et al., 2015 | 2013 | 2013-08-01 | Hyatt Resort Marina, Mission Bay, CA, California, USA | Non-native | 32.7634 | -117.2397 |
767518 | Ruiz et al., 2015 | 2013 | 2013-08-03 | Mission Bay Sport Center, Mission Bay, CA, California, USA | Non-native | 32.7857 | -117.2495 |
767534 | Ruiz et al., 2015 | 2013 | 2013-07-30 | Hilton Resort Docks, Mission Bay, CA, California, USA | Non-native | 32.7791 | -117.2128 |
767547 | Ruiz et al., 2015 | 2013 | 2013-08-02 | The Dana Marina, Mission Bay, CA, California, USA | Non-native | 32.7671 | -117.2363 |
767674 | Ruiz et al., 2015 | 2013 | 2013-07-17 | Naval Station San Diego, San Diego Bay, CA, California, USA | Non-native | 32.6867 | -117.1333 |
767690 | Ruiz et al., 2015 | 2013 | 2013-07-24 | NAB ACU-1 Docks, San Diego Bay, CA, California, USA | Non-native | 32.6786 | -117.1615 |
767714 | Ruiz et al., 2015 | 2013 | 2013-07-21 | Cabrillo Isle Marina, San Diego Bay, CA, California, USA | Non-native | 32.7272 | -117.1995 |
767727 | Ruiz et al., 2015 | 2013 | 2013-07-22 | Coronado Cays Marina, San Diego Bay, CA, California, USA | Non-native | 32.6257 | -117.1309 |
767740 | Ruiz et al., 2015 | 2013 | 2013-07-18 | NAB Fiddlers Cove, San Diego Bay, CA, California, USA | Non-native | 32.6524 | -117.1486 |
767755 | Ruiz et al., 2015 | 2013 | 2013-07-26 | Pier 32 Marina, San Diego Bay, CA, California, USA | Non-native | 32.6516 | -117.1077 |
767770 | Ruiz et al., 2015 | 2013 | 2013-07-20 | Chula Vista Marina, San Diego Bay, CA, California, USA | Non-native | 32.6252 | -117.1036 |
767780 | Ruiz et al., 2015 | 2013 | 2013-07-28 | Marriott Marquis and Marina, San Diego Bay, CA, California, USA | Non-native | 32.7059 | -117.1655 |
768557 | Ruiz et al., 2021b | 2017 | 2017-07-24 | Alamitos Bay Marina, San Pedro Bay, California, USA | Non-native | 33.7487 | -118.1136 |
768916 | Ruiz et al., 2021b | 2017 | 2017-07-29 | Pacific Yacht Landing, San Pedro Bay, California, USA | Non-native | 33.7654 | -118.2514 |
769085 | Ruiz et al., 2021b | 2017 | 2017-07-27 | Peter's Landing Marina, San Pedro Bay, California, USA | Non-native | 33.7256 | -118.0758 |
769314 | Ruiz et al., 2021b | 2018 | 2018-08-06 | Shoreline Public Marina, San Pedro Bay, California, USA | Non-native | 33.7586 | -118.1856 |
769322 | Ruiz et al., 2021b | 2018 | 2018-08-06 | Shoreline Public Marina, San Pedro Bay, California, USA | Non-native | 33.7586 | -118.1856 |
769340 | Ruiz et al., 2021b | 2018 | 2018-08-06 | Shoreline Public Marina, San Pedro Bay, California, USA | Non-native | 33.7586 | -118.1856 |
769366 | Ruiz et al., 2021b | 2018 | 2018-08-06 | Shoreline Public Marina, San Pedro Bay, California, USA | Non-native | 33.7586 | -118.1856 |
769378 | Ruiz et al., 2021b | 2018 | 2018-08-06 | Shoreline Public Marina, San Pedro Bay, California, USA | Non-native | 33.7586 | -118.1856 |
769384 | Ruiz et al., 2021b | 2018 | 2018-08-10 | Alamitos Bay Marina, San Pedro Bay, California, USA | Non-native | 33.7494 | -118.1154 |
769392 | Ruiz et al., 2021b | 2018 | 2018-08-10 | Alamitos Bay Marina, San Pedro Bay, California, USA | Non-native | 33.7494 | -118.1154 |
769404 | Ruiz et al., 2021b | 2018 | 2018-08-10 | Alamitos Bay Marina, San Pedro Bay, California, USA | Non-native | 33.7494 | -118.1154 |
769415 | Ruiz et al., 2021b | 2018 | 2018-08-10 | Alamitos Bay Marina, San Pedro Bay, California, USA | Non-native | 33.7494 | -118.1154 |
769424 | Ruiz et al., 2021b | 2018 | 2018-08-10 | Alamitos Bay Marina, San Pedro Bay, California, USA | Non-native | 33.7494 | -118.1154 |
769433 | Ruiz et al., 2021b | 2018 | 2018-08-10 | Alamitos Bay Marina, San Pedro Bay, California, USA | Non-native | 33.7494 | -118.1154 |
769444 | Ruiz et al., 2021b | 2018 | 2018-08-10 | Alamitos Bay Marina, San Pedro Bay, California, USA | Non-native | 33.7494 | -118.1154 |
769487 | Ruiz et al., 2021b | 2018 | 2018-08-12 | Pacific Yacht Landing, San Pedro Bay, California, USA | Non-native | 33.7654 | -118.2514 |
769497 | Ruiz et al., 2021b | 2018 | 2018-08-12 | Pacific Yacht Landing, San Pedro Bay, California, USA | Non-native | 33.7654 | -118.2514 |
769505 | Ruiz et al., 2021b | 2018 | 2018-08-12 | Pacific Yacht Landing, San Pedro Bay, California, USA | Non-native | 33.7654 | -118.2514 |
769528 | Ruiz et al., 2021b | 2018 | 2018-08-12 | Pacific Yacht Landing, San Pedro Bay, California, USA | Non-native | 33.7654 | -118.2514 |
769559 | Ruiz et al., 2021b | 2018 | 2018-08-12 | Pacific Yacht Landing, San Pedro Bay, California, USA | Non-native | 33.7654 | -118.2514 |
769569 | Ruiz et al., 2021b | 2018 | 2018-08-07 | Peter's Landing Marina, San Pedro Bay, California, USA | Non-native | 33.7252 | -118.0766 |
769584 | Ruiz et al., 2021b | 2018 | 2018-08-07 | Peter's Landing Marina, San Pedro Bay, California, USA | Non-native | 33.7252 | -118.0766 |
769594 | Ruiz et al., 2021b | 2018 | 2018-08-07 | Peter's Landing Marina, San Pedro Bay, California, USA | Non-native | 33.7252 | -118.0766 |
769605 | Ruiz et al., 2021b | 2018 | 2018-08-07 | Peter's Landing Marina, San Pedro Bay, California, USA | Non-native | 33.7252 | -118.0766 |
769614 | Ruiz et al., 2021b | 2018 | 2018-08-07 | Peter's Landing Marina, San Pedro Bay, California, USA | Non-native | 33.7252 | -118.0766 |
769627 | Ruiz et al., 2021b | 2018 | 2018-08-07 | Peter's Landing Marina, San Pedro Bay, California, USA | Non-native | 33.7252 | -118.0766 |
769638 | Ruiz et al., 2021b | 2018 | 2018-08-07 | Peter's Landing Marina, San Pedro Bay, California, USA | Non-native | 33.7252 | -118.0766 |
769656 | Ruiz et al., 2021b | 2018 | 2018-08-07 | Peter's Landing Marina, San Pedro Bay, California, USA | Non-native | 33.7252 | -118.0766 |
769771 | Ruiz et al., 2021b | 2018 | 2018-08-13 | Cabrillo Marina, San Pedro Bay, California, USA | Non-native | 33.7219 | -118.2779 |
769824 | Ruiz et al., 2021b | 2018 | 2018-08-13 | Cabrillo Marina, San Pedro Bay, California, USA | Non-native | 33.7219 | -118.2779 |
769856 | Ruiz et al., 2021b | 2018 | 2018-08-09 | Port of LB Pier D, San Pedro Bay, California, USA | Non-native | 33.7595 | -118.2174 |
769938 | Ruiz et al., 2021b | 2018 | 2018-08-08 | Port of LB Matson, San Pedro Bay, California, USA | Non-native | 33.7729 | -118.2178 |
769944 | Ruiz et al., 2021b | 2018 | 2018-08-08 | Port of LB Matson, San Pedro Bay, California, USA | Non-native | 33.7729 | -118.2178 |
769956 | Ruiz et al., 2021b | 2018 | 2018-08-08 | Port of LB Matson, San Pedro Bay, California, USA | Non-native | 33.7729 | -118.2178 |
769962 | Ruiz et al., 2021b | 2018 | 2018-08-08 | Port of LB Matson, San Pedro Bay, California, USA | Non-native | 33.7729 | -118.2178 |
769979 | Ruiz et al., 2021b | 2018 | 2018-08-08 | Port of LB Matson, San Pedro Bay, California, USA | Non-native | 33.7729 | -118.2178 |
769998 | Ruiz et al., 2021b | 2018 | 2018-08-08 | Port of LB Matson, San Pedro Bay, California, USA | Non-native | 33.7729 | -118.2178 |
770258 | Ruiz et al., 2021b | 2017 | 2017-08-03 | Balboa Bay Resort, Newport Bay, California, USA | Non-native | 33.6147 | -117.9138 |
770272 | Ruiz et al., 2021b | 2017 | 2017-08-03 | Balboa Bay Resort, Newport Bay, California, USA | Non-native | 33.6147 | -117.9138 |
770287 | Ruiz et al., 2021b | 2017 | 2017-08-03 | Balboa Bay Resort, Newport Bay, California, USA | Non-native | 33.6147 | -117.9138 |
770298 | Ruiz et al., 2021b | 2017 | 2017-08-03 | Balboa Bay Resort, Newport Bay, California, USA | Non-native | 33.6147 | -117.9138 |
770310 | Ruiz et al., 2021b | 2017 | 2017-08-03 | Balboa Bay Resort, Newport Bay, California, USA | Non-native | 33.6147 | -117.9138 |
770321 | Ruiz et al., 2021b | 2017 | 2017-08-07 | Balboa Yacht Basin A & B, Newport Bay, California, USA | Non-native | 33.6103 | -117.8949 |
770336 | Ruiz et al., 2021b | 2017 | 2017-08-07 | Balboa Yacht Basin A & B, Newport Bay, California, USA | Non-native | 33.6103 | -117.8949 |
770349 | Ruiz et al., 2021b | 2017 | 2017-08-07 | Balboa Yacht Basin A & B, Newport Bay, California, USA | Non-native | 33.6103 | -117.8949 |
770362 | Ruiz et al., 2021b | 2017 | 2017-08-07 | Balboa Yacht Basin A & B, Newport Bay, California, USA | Non-native | 33.6103 | -117.8949 |
770372 | Ruiz et al., 2021b | 2017 | 2017-08-07 | Balboa Yacht Basin A & B, Newport Bay, California, USA | Non-native | 33.6103 | -117.8949 |
770383 | Ruiz et al., 2021b | 2017 | 2017-08-07 | Balboa Yacht Basin D & E, Newport Bay, California, USA | Non-native | 33.6102 | -117.8967 |
770393 | Ruiz et al., 2021b | 2017 | 2017-08-07 | Balboa Yacht Basin D & E, Newport Bay, California, USA | Non-native | 33.6102 | -117.8967 |
770406 | Ruiz et al., 2021b | 2017 | 2017-08-07 | Balboa Yacht Basin D & E, Newport Bay, California, USA | Non-native | 33.6102 | -117.8967 |
770434 | Ruiz et al., 2021b | 2017 | 2017-08-06 | Harbor Patrol Public Dock, Newport Bay, California, USA | Non-native | 33.6023 | -117.8835 |
770442 | Ruiz et al., 2021b | 2017 | 2017-08-06 | Harbor Patrol Public Dock, Newport Bay, California, USA | Non-native | 33.6023 | -117.8835 |
770453 | Ruiz et al., 2021b | 2017 | 2017-08-06 | Harbor Patrol Public Dock, Newport Bay, California, USA | Non-native | 33.6023 | -117.8835 |
770463 | Ruiz et al., 2021b | 2017 | 2017-08-06 | Harbor Patrol Public Dock, Newport Bay, California, USA | Non-native | 33.6023 | -117.8835 |
770471 | Ruiz et al., 2021b | 2017 | 2017-08-06 | Harbor Patrol Public Dock, Newport Bay, California, USA | Non-native | 33.6023 | -117.8835 |
770486 | Ruiz et al., 2021b | 2017 | 2017-08-05 | Harbor Tower Marina, Newport Bay, California, USA | Non-native | 33.6203 | -117.9279 |
770495 | Ruiz et al., 2021b | 2017 | 2017-08-05 | Harbor Tower Marina, Newport Bay, California, USA | Non-native | 33.6203 | -117.9279 |
770512 | Ruiz et al., 2021b | 2017 | 2017-08-05 | Harbor Tower Marina, Newport Bay, California, USA | Non-native | 33.6203 | -117.9279 |
770523 | Ruiz et al., 2021b | 2017 | 2017-08-04 | Hill's Boat Service, Newport Bay, California, USA | Non-native | 33.6026 | -117.8984 |
770560 | Ruiz et al., 2021b | 2017 | 2017-08-04 | Hill's Boat Service, Newport Bay, California, USA | Non-native | 33.6026 | -117.8984 |
770571 | Ruiz et al., 2021b | 2017 | 2017-08-04 | Hill's Boat Service, Newport Bay, California, USA | Non-native | 33.6026 | -117.8984 |
770583 | Ruiz et al., 2021b | 2017 | 2017-08-04 | Marina Park, Newport Bay, California, USA | Non-native | 33.6085 | -117.9222 |
770603 | Ruiz et al., 2021b | 2017 | 2017-08-04 | Marina Park, Newport Bay, California, USA | Non-native | 33.6085 | -117.9222 |
770613 | Ruiz et al., 2021b | 2017 | 2017-08-04 | Marina Park, Newport Bay, California, USA | Non-native | 33.6085 | -117.9222 |
770634 | Ruiz et al., 2021b | 2017 | 2017-08-15 | Newport Dunes, Newport Bay, California, USA | Non-native | 33.6193 | -117.8959 |
770646 | Ruiz et al., 2021b | 2017 | 2017-08-15 | Newport Dunes, Newport Bay, California, USA | Non-native | 33.6193 | -117.8959 |
770655 | Ruiz et al., 2021b | 2017 | 2017-08-15 | Newport Dunes, Newport Bay, California, USA | Non-native | 33.6193 | -117.8959 |
770664 | Ruiz et al., 2021b | 2017 | 2017-08-15 | Newport Dunes, Newport Bay, California, USA | Non-native | 33.6193 | -117.8959 |
770675 | Ruiz et al., 2021b | 2017 | 2017-08-15 | Newport Dunes, Newport Bay, California, USA | Non-native | 33.6193 | -117.8959 |
770690 | Ruiz et al., 2021b | 2017 | 2017-08-03 | Rhine Wharf Public Pier, Newport Bay, California, USA | Non-native | 33.6148 | -117.9266 |
770697 | Ruiz et al., 2021b | 2017 | 2017-08-03 | Rhine Wharf Public Pier, Newport Bay, California, USA | Non-native | 33.6148 | -117.9266 |
770706 | Ruiz et al., 2021b | 2017 | 2017-08-03 | Rhine Wharf Public Pier, Newport Bay, California, USA | Non-native | 33.6148 | -117.9266 |
770715 | Ruiz et al., 2021b | 2017 | 2017-08-03 | Rhine Wharf Public Pier, Newport Bay, California, USA | Non-native | 33.6148 | -117.9266 |
770722 | Ruiz et al., 2021b | 2017 | 2017-08-03 | Rhine Wharf Public Pier, Newport Bay, California, USA | Non-native | 33.6148 | -117.9266 |
776305 | Ruiz et al., 2022 | 2014 | 2014-09-12 | Ballena Isle Marina, San Francisco Bay, California, USA | Non-native | 37.7662 | -122.2656 |
776306 | Ruiz et al., 2022 | 2014 | 2014-09-12 | Ballena Isle Marina, San Francisco Bay, California, USA | Non-native | 37.7662 | -122.2656 |
776307 | Ruiz et al., 2022 | 2014 | 2014-09-12 | Ballena Isle Marina, San Francisco Bay, California, USA | Non-native | 37.7662 | -122.2656 |
776308 | Ruiz et al., 2022 | 2014 | 2014-09-12 | Ballena Isle Marina, San Francisco Bay, California, USA | Non-native | 37.7662 | -122.2656 |
776309 | Ruiz et al., 2022 | 2014 | 2014-09-18 | Oakland Yacht Club, San Francisco Bay, California, USA | Non-native | 37.7835 | -122.2630 |
776310 | Ruiz et al., 2022 | 2014 | 2014-09-18 | Oakland Yacht Club, San Francisco Bay, California, USA | Non-native | 37.7835 | -122.2630 |
776311 | Ruiz et al., 2022 | 2014 | 2014-09-18 | Oakland Yacht Club, San Francisco Bay, California, USA | Non-native | 37.7835 | -122.2630 |
776312 | Ruiz et al., 2022 | 2014 | 2014-09-18 | Oakland Yacht Club, San Francisco Bay, California, USA | Non-native | 37.7835 | -122.2630 |
776313 | Ruiz et al., 2022 | 2014 | 2014-09-18 | Oakland Yacht Club, San Francisco Bay, California, USA | Non-native | 37.7835 | -122.2630 |
776314 | Ruiz et al., 2022 | 2014 | 2014-09-18 | Oakland Yacht Club, San Francisco Bay, California, USA | Non-native | 37.7835 | -122.2630 |
776315 | Ruiz et al., 2022 | 2014 | 2014-09-18 | Oakland Yacht Club, San Francisco Bay, California, USA | Non-native | 37.7835 | -122.2630 |
776316 | Ruiz et al., 2022 | 2014 | 2014-09-18 | Oakland Yacht Club, San Francisco Bay, California, USA | Non-native | 37.7835 | -122.2630 |
776317 | Ruiz et al., 2022 | 2014 | 2014-09-18 | Oakland Yacht Club, San Francisco Bay, California, USA | Non-native | 37.7835 | -122.2630 |
776318 | Ruiz et al., 2022 | 2015 | 2015-09-24 | Oakland Yacht Club, San Francisco Bay, California, USA | Non-native | 37.7839 | -122.2641 |
776319 | Ruiz et al., 2022 | 2015 | 2015-09-24 | Oakland Yacht Club, San Francisco Bay, California, USA | Non-native | 37.7839 | -122.2641 |
776320 | Ruiz et al., 2022 | 2015 | 2015-09-24 | Oakland Yacht Club, San Francisco Bay, California, USA | Non-native | 37.7839 | -122.2641 |
776321 | Ruiz et al., 2022 | 2015 | 2015-09-24 | Oakland Yacht Club, San Francisco Bay, California, USA | Non-native | 37.7839 | -122.2641 |
776322 | Ruiz et al., 2022 | 2015 | 2015-09-24 | Oakland Yacht Club, San Francisco Bay, California, USA | Non-native | 37.7839 | -122.2641 |
776323 | Ruiz et al., 2022 | 2015 | 2015-09-24 | Oakland Yacht Club, San Francisco Bay, California, USA | Non-native | 37.7839 | -122.2641 |
776324 | Ruiz et al., 2022 | 2015 | 2015-09-24 | Oakland Yacht Club, San Francisco Bay, California, USA | Non-native | 37.7839 | -122.2641 |
776325 | Ruiz et al., 2022 | 2015 | 2015-09-24 | Oakland Yacht Club, San Francisco Bay, California, USA | Non-native | 37.7839 | -122.2641 |
776326 | Ruiz et al., 2022 | 2015 | 2015-09-24 | Oakland Yacht Club, San Francisco Bay, California, USA | Non-native | 37.7839 | -122.2641 |
776327 | Ruiz et al., 2022 | 2015 | 2015-09-21 | Redwood City Marina, San Francisco Bay, California, USA | Non-native | 37.5022 | -122.2129 |
776328 | Ruiz et al., 2022 | 2015 | 2015-09-21 | Redwood City Marina, San Francisco Bay, California, USA | Non-native | 37.5022 | -122.2129 |
776329 | Ruiz et al., 2022 | 2015 | 2015-09-21 | Redwood City Marina, San Francisco Bay, California, USA | Non-native | 37.5022 | -122.2129 |
776330 | Ruiz et al., 2022 | 2015 | 2015-09-21 | Redwood City Marina, San Francisco Bay, California, USA | Non-native | 37.5022 | -122.2129 |
776331 | Ruiz et al., 2022 | 2015 | 2015-09-21 | Redwood City Marina, San Francisco Bay, California, USA | Non-native | 37.5022 | -122.2129 |
776332 | Ruiz et al., 2022 | 2015 | 2015-09-21 | Redwood City Marina, San Francisco Bay, California, USA | Non-native | 37.5022 | -122.2129 |
776333 | Ruiz et al., 2022 | 2015 | 2015-09-21 | Redwood City Marina, San Francisco Bay, California, USA | Non-native | 37.5022 | -122.2129 |
776334 | Ruiz et al., 2022 | 2015 | 2015-09-21 | Redwood City Marina, San Francisco Bay, California, USA | Non-native | 37.5022 | -122.2129 |
776335 | Ruiz et al., 2022 | 2015 | 2015-09-21 | Redwood City Marina, San Francisco Bay, California, USA | Non-native | 37.5022 | -122.2129 |
776336 | Ruiz et al., 2022 | 2015 | 2015-09-21 | Redwood City Marina, San Francisco Bay, California, USA | Non-native | 37.5022 | -122.2129 |
776337 | Ruiz et al., 2022 | 2015 | 2015-09-14 | Richmond Marina Bay Yacht Harbor, San Francisco Bay, California, USA | Non-native | 37.9129 | -122.3492 |
776338 | Ruiz et al., 2022 | 2015 | 2015-09-14 | Richmond Marina Bay Yacht Harbor, San Francisco Bay, California, USA | Non-native | 37.9129 | -122.3492 |
776339 | Ruiz et al., 2022 | 2015 | 2015-09-14 | Richmond Marina Bay Yacht Harbor, San Francisco Bay, California, USA | Non-native | 37.9129 | -122.3492 |
776340 | Ruiz et al., 2022 | 2015 | 2015-09-14 | Richmond Marina Bay Yacht Harbor, San Francisco Bay, California, USA | Non-native | 37.9129 | -122.3492 |
776341 | Ruiz et al., 2022 | 2015 | 2015-09-14 | Richmond Marina Bay Yacht Harbor, San Francisco Bay, California, USA | Non-native | 37.9129 | -122.3492 |
776342 | Ruiz et al., 2022 | 2015 | 2015-09-14 | Richmond Marina Bay Yacht Harbor, San Francisco Bay, California, USA | Non-native | 37.9129 | -122.3492 |
776343 | Ruiz et al., 2022 | 2015 | 2015-09-17 | Ballena Isle Marina, San Francisco Bay, California, USA | Non-native | 37.7679 | -122.2863 |
776344 | Ruiz et al., 2022 | 2015 | 2015-09-17 | Ballena Isle Marina, San Francisco Bay, California, USA | Non-native | 37.7679 | -122.2863 |
776345 | Ruiz et al., 2022 | 2015 | 2015-09-17 | Ballena Isle Marina, San Francisco Bay, California, USA | Non-native | 37.7679 | -122.2863 |
776346 | Ruiz et al., 2022 | 2015 | 2015-09-17 | Ballena Isle Marina, San Francisco Bay, California, USA | Non-native | 37.7679 | -122.2863 |
776347 | Ruiz et al., 2022 | 2015 | 2015-09-17 | Ballena Isle Marina, San Francisco Bay, California, USA | Non-native | 37.7679 | -122.2863 |
776348 | Ruiz et al., 2022 | 2015 | 2015-09-17 | Ballena Isle Marina, San Francisco Bay, California, USA | Non-native | 37.7679 | -122.2863 |
776349 | Ruiz et al., 2022 | 2015 | 2015-09-17 | Ballena Isle Marina, San Francisco Bay, California, USA | Non-native | 37.7679 | -122.2863 |
776350 | Ruiz et al., 2022 | 2015 | 2015-09-17 | Ballena Isle Marina, San Francisco Bay, California, USA | Non-native | 37.7679 | -122.2863 |
776351 | Ruiz et al., 2022 | 2015 | 2015-09-17 | Ballena Isle Marina, San Francisco Bay, California, USA | Non-native | 37.7679 | -122.2863 |
776352 | Ruiz et al., 2022 | 2015 | 2015-09-17 | Ballena Isle Marina, San Francisco Bay, California, USA | Non-native | 37.7679 | -122.2863 |
776353 | Ruiz et al., 2022 | 2015 | 2015-09-22 | Coyote Point Marina, San Francisco Bay, California, USA | Non-native | 37.5887 | -122.3164 |
776354 | Ruiz et al., 2022 | 2015 | 2015-09-22 | Coyote Point Marina, San Francisco Bay, California, USA | Non-native | 37.5887 | -122.3164 |
776355 | Ruiz et al., 2022 | 2015 | 2015-09-22 | Coyote Point Marina, San Francisco Bay, California, USA | Non-native | 37.5887 | -122.3164 |
776356 | Ruiz et al., 2022 | 2015 | 2015-09-22 | Coyote Point Marina, San Francisco Bay, California, USA | Non-native | 37.5887 | -122.3164 |
776357 | Ruiz et al., 2022 | 2015 | 2015-09-22 | Coyote Point Marina, San Francisco Bay, California, USA | Non-native | 37.5887 | -122.3164 |
776358 | Ruiz et al., 2022 | 2015 | 2015-09-22 | Coyote Point Marina, San Francisco Bay, California, USA | Non-native | 37.5887 | -122.3164 |
776359 | Ruiz et al., 2022 | 2015 | 2015-09-22 | Coyote Point Marina, San Francisco Bay, California, USA | Non-native | 37.5887 | -122.3164 |
776360 | Ruiz et al., 2022 | 2015 | 2015-09-16 | San Leandro Marina, San Francisco Bay, California, USA | Non-native | 37.6978 | -122.1928 |
776361 | Ruiz et al., 2022 | 2015 | 2015-09-16 | San Leandro Marina, San Francisco Bay, California, USA | Non-native | 37.6978 | -122.1928 |
776362 | Ruiz et al., 2022 | 2015 | 2015-09-16 | San Leandro Marina, San Francisco Bay, California, USA | Non-native | 37.6978 | -122.1928 |
776363 | Ruiz et al., 2022 | 2015 | 2015-09-16 | San Leandro Marina, San Francisco Bay, California, USA | Non-native | 37.6978 | -122.1928 |
776364 | Ruiz et al., 2022 | 2015 | 2015-09-16 | San Leandro Marina, San Francisco Bay, California, USA | Non-native | 37.6978 | -122.1928 |
776365 | Ruiz et al., 2022 | 2015 | 2015-09-16 | San Leandro Marina, San Francisco Bay, California, USA | Non-native | 37.6978 | -122.1928 |
776366 | Ruiz et al., 2022 | 2015 | 2015-09-16 | San Leandro Marina, San Francisco Bay, California, USA | Non-native | 37.6978 | -122.1928 |
776367 | Ruiz et al., 2022 | 2015 | 2015-09-16 | San Leandro Marina, San Francisco Bay, California, USA | Non-native | 37.6978 | -122.1928 |
776368 | Ruiz et al., 2022 | 2016 | 2016-09-22 | Coyote Point Marina, San Francisco Bay, California, USA | Non-native | 37.5898 | -122.3165 |
776369 | Ruiz et al., 2022 | 2016 | 2016-09-22 | Coyote Point Marina, San Francisco Bay, California, USA | Non-native | 37.5898 | -122.3165 |
776370 | Ruiz et al., 2022 | 2016 | 2016-09-22 | Coyote Point Marina, San Francisco Bay, California, USA | Non-native | 37.5898 | -122.3165 |
776371 | Ruiz et al., 2022 | 2016 | 2016-09-22 | Coyote Point Marina, San Francisco Bay, California, USA | Non-native | 37.5898 | -122.3165 |
776372 | Ruiz et al., 2022 | 2016 | 2016-09-22 | Coyote Point Marina, San Francisco Bay, California, USA | Non-native | 37.5898 | -122.3165 |
776373 | Ruiz et al., 2022 | 2016 | 2016-09-22 | Coyote Point Marina, San Francisco Bay, California, USA | Non-native | 37.5898 | -122.3165 |
776374 | Ruiz et al., 2022 | 2016 | 2016-09-22 | Coyote Point Marina, San Francisco Bay, California, USA | Non-native | 37.5898 | -122.3165 |
776375 | Ruiz et al., 2022 | 2016 | 2016-09-22 | Coyote Point Marina, San Francisco Bay, California, USA | Non-native | 37.5898 | -122.3165 |
776376 | Ruiz et al., 2022 | 2016 | 2016-09-26 | Redwood City Marina, San Francisco Bay, California, USA | Non-native | 37.5027 | -122.2138 |
776377 | Ruiz et al., 2022 | 2016 | 2016-09-26 | Redwood City Marina, San Francisco Bay, California, USA | Non-native | 37.5027 | -122.2138 |
776378 | Ruiz et al., 2022 | 2016 | 2016-09-26 | Redwood City Marina, San Francisco Bay, California, USA | Non-native | 37.5027 | -122.2138 |
776379 | Ruiz et al., 2022 | 2016 | 2016-09-26 | Redwood City Marina, San Francisco Bay, California, USA | Non-native | 37.5027 | -122.2138 |
776380 | Ruiz et al., 2022 | 2016 | 2016-09-26 | Redwood City Marina, San Francisco Bay, California, USA | Non-native | 37.5027 | -122.2138 |
776381 | Ruiz et al., 2022 | 2016 | 2016-09-26 | Redwood City Marina, San Francisco Bay, California, USA | Non-native | 37.5027 | -122.2138 |
776382 | Ruiz et al., 2022 | 2016 | 2016-09-26 | Redwood City Marina, San Francisco Bay, California, USA | Non-native | 37.5027 | -122.2138 |
776383 | Ruiz et al., 2022 | 2016 | 2016-09-26 | Redwood City Marina, San Francisco Bay, California, USA | Non-native | 37.5027 | -122.2138 |
776384 | Ruiz et al., 2022 | 2016 | 2016-09-12 | Richmond Marina Bay Yacht Harbor, San Francisco Bay, California, USA | Non-native | 37.9133 | -122.3500 |
776385 | Ruiz et al., 2022 | 2016 | 2016-09-12 | Richmond Marina Bay Yacht Harbor, San Francisco Bay, California, USA | Non-native | 37.9133 | -122.3500 |
776386 | Ruiz et al., 2022 | 2016 | 2016-09-12 | Richmond Marina Bay Yacht Harbor, San Francisco Bay, California, USA | Non-native | 37.9133 | -122.3500 |
776387 | Ruiz et al., 2022 | 2016 | 2016-09-13 | Ballena Isle Marina, San Francisco Bay, California, USA | Non-native | 37.7679 | -122.2863 |
776388 | Ruiz et al., 2022 | 2016 | 2016-09-13 | Ballena Isle Marina, San Francisco Bay, California, USA | Non-native | 37.7679 | -122.2863 |
776389 | Ruiz et al., 2022 | 2016 | 2016-09-13 | Ballena Isle Marina, San Francisco Bay, California, USA | Non-native | 37.7679 | -122.2863 |
776390 | Ruiz et al., 2022 | 2016 | 2016-09-13 | Ballena Isle Marina, San Francisco Bay, California, USA | Non-native | 37.7679 | -122.2863 |
776391 | Ruiz et al., 2022 | 2016 | 2016-09-27 | Oakland Yacht Club, San Francisco Bay, California, USA | Non-native | 37.7831 | -122.2630 |
776392 | Ruiz et al., 2022 | 2016 | 2016-09-27 | Oakland Yacht Club, San Francisco Bay, California, USA | Non-native | 37.7831 | -122.2630 |
776393 | Ruiz et al., 2022 | 2016 | 2016-09-27 | Oakland Yacht Club, San Francisco Bay, California, USA | Non-native | 37.7831 | -122.2630 |
776394 | Ruiz et al., 2022 | 2016 | 2016-09-27 | Oakland Yacht Club, San Francisco Bay, California, USA | Non-native | 37.7831 | -122.2630 |
776395 | Ruiz et al., 2022 | 2016 | 2016-09-27 | Oakland Yacht Club, San Francisco Bay, California, USA | Non-native | 37.7831 | -122.2630 |
776396 | Ruiz et al., 2022 | 2016 | 2016-09-27 | Oakland Yacht Club, San Francisco Bay, California, USA | Non-native | 37.7831 | -122.2630 |
776397 | Ruiz et al., 2022 | 2016 | 2016-09-27 | Oakland Yacht Club, San Francisco Bay, California, USA | Non-native | 37.7831 | -122.2630 |
776398 | Ruiz et al., 2022 | 2016 | 2016-09-14 | San Leandro Marina, San Francisco Bay, California, USA | Non-native | 37.6978 | -122.1928 |
776399 | Ruiz et al., 2022 | 2016 | 2016-09-14 | San Leandro Marina, San Francisco Bay, California, USA | Non-native | 37.6978 | -122.1928 |
776400 | Ruiz et al., 2022 | 2016 | 2016-09-14 | San Leandro Marina, San Francisco Bay, California, USA | Non-native | 37.6978 | -122.1928 |
776401 | Ruiz et al., 2022 | 2016 | 2016-09-14 | San Leandro Marina, San Francisco Bay, California, USA | Non-native | 37.6978 | -122.1928 |
References
Abdel-Salam, Kh. M.; Ramadan, Sh. E. (2008) Fouling Bryozoa from some Alexandria harbours, Egypt. (II) Encrusting species, Mediterranean Marine Science 9(2): 5-20Abdelsalam, Khaled Mahmood (2018) First record of the exotic lysmatid shrimp Lysmata vittata (Stimpson, 1860) (Decapoda: Caridea: Lysmatidae) from the Egyptian Mediterranean coast, Mediterranean Marine Science 19(1): 124-131
Alvarez-Leon, R. ; Banta, W. C. (1984) [Bryozoans of three estuaries adjacent to Mazatlan, northeast coast of Mexico], Studies on Neotropical Fauna and Environment 19(4): 209-218
Amat, Jaen Nieto; Tempera, Fernando (2009) Zoobotryon verticillatum Della Chiaje, 1822 (Bryozoa), a new occurrence in the archipelago of the Azores (North-Eastern Atlantic), Marine Pollution Bulletin 58: 761-764
Antit, M.; Gofas, S.; Salas, C.; Azzouna, A. (2011) One hundred years after Pinctada: an update on alien Mollusca in Tunisia, Mediterranean Marine Science 12(1): 53-73
Armstrong, John D.; Bean, Colin W.; Wells, Alan (2018) The Scottish invasion of pink salmon in 2017, Journal of Fish Biology 93: 8-11
DOI: 10.1111/jfb.13680
Asakura, A. (1992) Recent introductions of marine benthos into Tokyo Bay (review): Process of invasion into an urban ecosystem with discussion on the factors inducing their successful introduction, Journal of the Natural History Museum and Institute, Chiba 2(1): 1-14
Baker, H. R. (1984) Diversity and zoogeography of marine Tubificidae (Annelida, Oligochaeta), with notes on variation in widespread species, Hydrobiologia 115: 191-196
Barnes, Robert D. (1983) Invertebrate Zoology, Saunders, Philadelphia. Pp. 883
Beshai, Ryan A.; Truong, Danny A.; Henry, Amy K. Sorte, Cascade J. B. (2022) Biotic resistance or invasional meltdown? Diversity reduces invasibility but not exotic dominance in southern California epibenthic communities, Biological Invasions 25(2): 533-549
https://doi.org/10.1007/s10530-022-02932-1
Brock, Brian J. (1985) Bryozoa: Ordovician to Recent, Olsen & Olsen, Fredensborg. Pp. 45-49
Califonria Department of Fisheries and Wildlife 2021 Invasive Algae Response (Aulerpa spp.). <missing URL>
Canning-Clode, João; Fofonoff, Paul; McCann, Linda; Carlton, James T.; Ruiz, Gregory (2013) Marine invasions on a subtropical island: fouling studies and new records in a recent marina on Madeira Island (Eastern Atlantic Ocean), Aquatic Invasions 8(3): 261-270
Carlton, James T.; Eldredge, Lucius (2009) Marine bioinvasions of Hawaii: The introduced and cryptogenic marine and estuarine animals and plants of the Hawaiian archipelago., Bishop Museum Bulletin in Cultural and Environmental Studies 4: 1-202
Carlton, James T.; Ruckelshaus, Mary H. (1997) Nonindigenous marine invertebrates and algae of Florida, In: Simberloff, Daniel, Schmitz, Don C., Brown, Tom C.(Eds.) Strangers in Paradise: Impact and Management of Nonindigenous Species in Florida. , Washington, D.C.. Pp. 187-201
Çinar, Melih Ertan and 7 authors (2021) Current status (as of end of 2020) of marine alien species in Turkey, PLOS ONE 16: Published online
Cohen, Andrew N. and 12 authors (2002) Project report for the Southern California exotics expedition 2000: a rapid assessment survey of exotic species in sheltered coastal waters., In: (Eds.) . , Sacramento CA. Pp. 1-23
Cohen, Andrew N.; Carlton, James T. (1995) Nonindigenous aquatic species in a United States estuary: a case study of the biological invasions of the San Francisco Bay and Delta, U.S. Fish and Wildlife Service and National Sea Grant College Program (Connecticut Sea Grant), Washington DC, Silver Spring MD.. Pp. <missing location>
Coles S. L., DeFelice R. C., Eldredge, L. G. (2002b) Nonindigenous marine species at Waikîkî and Hawai`i kai, Oahu, Hawai`i, Bishop Museum Technical Report 25: 1-255
Coles, S. L.; DeFelice, R. C. : Eldredge, L. G. (2002a) Nonindigenous marine species in Kaneohe Bay, Oahu, Hawai`i, Bishop Museum Technical Report 24: 1-364
Coles, S. L.; Eldredge, L. G.; Kandel, F.; Reath, P. R.; Longenecker, K. (2004) <missing title>, Bishop Museum, Hawai‘i Biological Survey, Honolulu. Pp. <missing location>
Coles, S. L.; Reath, P. R.; Longenecker, K.; Bolick, Holly; Eldredge, L. G. (2004) <missing title>, Hawai‘i Community Foundation and the U. S. Fish and Wildlife Service, Honolulu. Pp. 1-187
Cook, Patricia L. (1968) Bryozoa (Polyzoa) from the coasts of tropical West Africa, Atlantide - Report 10: 115-262
Corriero, G.; Longo, C.; Mercurio, M.; Marchini, A.; Occhipinti-Ambrogi, A. (2007) Porifera and Bryozoa on artificial hard bottoms in the Venice Lagoon: Spatial distribution and temporal changes in the northern basin, Italian Journal of Zoology 74(1): 21-29
Corsini-Foka, Maria and 33 authors (2015) Inventory of alien and cryptogenic species of the Dodecanese (Aegean Sea, Greece): collaboration through COST action training school, Aquatic Invasions 10: In press
Cranfield, H.J.; Gordon, D.P.; Willan, R.C.; Marshall, B.A; Battershill, C.N.; Francis, M.P.; Nelson, W.A.; Glasby, C.J.; Read, G.B. (1998) <missing title>, The National Institute of Water and Atmospheric Research, New Zealand. Pp. <missing location>
Creary, Marcia M. (2003) A simplified field guide to the bryozoan species found on the roots of the red mangrove (Rhizophora mangle) in and around Kingston Harbour, Jamaica, W.I., Bulletin of Marine Science 73(2): 521-526
Crivellaro, Marcelo Schuler; Candido, Davi Volney; ilveira, Thiago Cesar Lima; Fonseca, Adriana Carvalhal; Segal, Barbara ´ (2022) A tool for a race against time: Dispersal simulations to support ongoing monitoring program of the invasive coral Tubastraea coccinea, Marine Pollution Bulletin 185(114354): Published online
https://doi.org/10.1016/j.marpolbul.2022.114354
D'Hondt, Jean-Loup (1975) Sur quelques bryozoaires de la region de Dakar, Bulletin de l nstitut Fondamental dAfrique Noire 37(1): 619-628
de Rivera, Catherine, and 27 authors (2005) Broad-scale non-indigenous species monitoring along the West Coast in National Marine Sanctuaries and National Estuarine Research Reserves report to National Fish and Wildlife Foundation, National Fish and Wildlife Foundation, Washington, D.C.. Pp. <missing location>
Fairey, Russell; Dunn, Roslyn; Sigala, Marco; Oliver, John (2002) Introduced aquatic species in California's coastal waters: Final Report, California Department of Fish and Game, Sacramento. Pp. <missing location>
Farrapeira, Cristiane Maria Rocha (2011) The introduction of the bryozoan Zoobotryon verticillatum (Della Chiaje, 1822) in northeast of Brazil: a cause for concern, Biological Invasions 13: 13-16
Fell, Paul E. (1975) Salinity tolerance and dessication reisitance of the brackish water sponge Haliclona loosanoffi, Journal of Experimental Zoology 194: 409-412
Galil, Bella S.; Gevili, Roy (2014) Zoobotryon verticillatum (Bryozoa: Ctenostomatida: Vesiculariidae), a new occurrence on the Mediterranean coast of Israel, Marine Biodiversity Records 7: e17
Gordon, Dennis P.; Mawatari, S.F. (1992) Atlas of marine-fouling bryozoa of New Zealand ports and harbors., Miscellaneous Publications of the New Zealand Oceanographic Institute 107: 1-52
Gossett, Lisa; Lester, Jim; Gonzalez, Lisa (2004) <missing title>, Environmental Institute of Houston, University of Houston, Houston TX. Pp. <missing location>
Gravier-Bonnet, Nicole; Migotto, Alvaro E. (2000) Gonangium development and medusoid of Nemalecium lighti (Hargitt, 1924) (Cnidaria: Hydrozoa, Haleciidae), Scientia Marina 64(Suppl. 1): 207-213
Guerra-García, J. M. and 7 authors (2011) Geographical expansion of the invader Caprella scaura (Crustacea: Amphipoda: Caprellidae) to the East Atlantic coast, Marine Biology 158: 2617-2622
Hewitt, C.L.; Campbell, M.L.; Thresher, R.E.; Martin, R.B. (1999) Marine Biological Invasions of Port Phillip Bay, Victoria, In: (Eds.) . , Hobart, Tasmania. Pp. <missing location>
Hewitt, Chad L. and 14 authors. (2004) Introduced and cryptogenic species in Port Phillip Bay, Victoria, Australia., Marine Biology 144: 183-202
Huang, Zongguo (Ed.), Junda Lin (Translator) (2001) Marine Species and Their Distributions in China's Seas, Krieger, Malabar, FL. Pp. <missing location>
Huisman, John M.; Jones, Diana S.; Wells, Fred E.; Burton, Timothy S. (2008) Introduced marine biota in Western Australian waters., Records of the Western Australian Museum 25: 1-44
Humara-Gil, Karla J.; Cruz-Gil, Christopher (2019) First record of the non-indigenous bryozoan Amathia verticillata (delle Chiaje, 1822) (Bryozoa, Vesiculariidae) in the southern Mexican Pacific, Check List 15(3): 515-522
Hyman, Libbie H. (1959) <missing title>, McGraw-Hill, New York. Pp. 350-408
Je, J-G.; Hong, J-S.; Yi, S-K. (1988) A study on the fouling organisms in the pearl oyster culture grounds in the southern coast of Korea., Ocean Research 10(1): 85-105.
Jebakumar, Jebarathnam Prince Prakash, and 5 authors (2017) The bryozoan Amathia verticillata(delle Chiaje, 1822) fouling harbours of the southeast coast of India: re-evaluating its status, BioInvasions Records 6(3): 214-216
Johnson, Leigh Taylor; Gonzalez, Jamie; Himes, Amber (2006) <missing title>, University of California Cooperative Extension – Sea Grant Extension Program, San Diego. Pp. 3
Kaminas, Alexandros; Shokouros-Oskarsson. Maria; Minasidis, Vasileios; Langeneck, Joachim; Kleitou, Periklis; Tiralongo; Crocetta, Fabio (2033) Filling gaps via citizen science: Phyllorhiza punctata von Lendenfeld, 1884 (Cnidaria: Scyphozoa: Mastigiidae) in Cyprus (eastern Mediterranean Sea), Bioinvasion Records 11(In press): In press
Kelmo, Francisco; Attrill, Martin J.; Gomes, Rilza C.T.; Jones, Malcolm B. (2004) El Nino induced local extinction of coral reef bryozoan species from Northern Bahia, Brazil, Biological Conservation 118: 609-617
Keough, M. J. ; Ross, J. (1999) Introduced fouling species in Port Phillip Bay., In: Hewitt, C. L.; Campbell; M.;Thresher, R.; Martin,(Eds.) Marine Biological Invasions of Port Phillip Bay, Victoria. , Hobart, Tasmania. Pp. 193-225
Knapp, I. S.; Godwin, L. S.; Smith, J. E.; Williams, C. J.; Bell, J. J. (2011) Records of non-indigenous marine species at Palmyra Atoll in the US Line Islands, Marine Biodiversity Records 4(e30): 1-7
Kourkoutmani, Polyxeni; Michaloudi, Evangelia (2022) First record of the calanoid copepod Pseudodiaptomus marinus Sato, 1913 in the North Aegean Sea, in Thessaloniki Bay, Greece, None 11(BioInvasions Re): Published online
Lambert, Gretchen (2019) Fouling ascidians (Chordata: Ascidiacea) of the Galápagos: Santa Cruz and Baltra Islands, Aquatic Invasions 14: 132-149
Lavigne, Andrea S. Sunesen, Ines Sar, Eugenia A. (2015) Morphological, taxonomic and nomenclatural analysis of species of Odontella, Trieres and Zygoceros (Triceratiaceae, Bacillariophyta) from Anegada Bay (Province of Buenos Aires, Argentina), Diatom Research 30: 307-331
Leca, L.; d'Hondt, J.-L. (1993) [Hippopetraliella tahitiensis n. sp. n. sp., a new Cheilostome bryozoan (Petraliellidae) from French Polynesia], Cahiers de Biologie Marine 34: 401-209
Madhaven Pillai, S. R. (1981) Further report on the taxonomy of fouling bryozoans of Bombay Harbour and vicinity., Journal of the Bombay Natural History Society 78: 317-329
Marchini, Agnese; Cunha, Maria R.; Occhipinti-Ambrogi, Anna (2007) First observations on bryozoans and entoprocts in the Ria de Aveiro (NW Portugal) including the first record of the Pacific invasive cheilostome Tricellaria inopinata., Marine Ecology 28(Suppl. 1): 154-160
Marchini, Agnese; Ferrario, Jasmine; Minchin, Dan (2015) Marinas may act as hubs for the spread of the pseudo-indigenous bryozoan Amathia verticillata (Delle Chiaje, 1822) and its associates, Scientia Marina 79(3): 355-365
Marcus, Ernst (1937) Bryozoarios Marinhos Brasileiros, Boletim de faculdade de filosofia, Ciencias e Letras, Universidade de sao Paulo, Zoologia 1: 1-224
Marshall, J. I.; Rowe, F. W. E.; Fisher; Smith, D. F. (1980) Alterations to the relative species-abundance of ascidians and barnacles in a fouling community due to screens, Australian Journal of Marine and Freshwater Research 31: <missing location>
Maturo, Frank J. S., Jr. (1957) A study of the Bryozoa of Beaufort, North Carolina and vicinity, Journal of the Elisha Mitchell Scientific Society 72(1): 11-68
Maturo, Frank J. S., Jr.; Schopf Thomas J. M. (1968) Ectoproct and entoproct type material: reexamination of species from New England and Bermuda named by A.E. Verrill, J.W. Dawson, and E. Desor, Postilla 120: 1-95
McCann, Linda D.;; McCuller, Megan I., Carlton, James T.[ ,Keith, Inti; Geller, Jonathan B.; Ruiz, Gregory M. (2019) Bryozoa (Cheilostomata, Ctenostomata, and Cyclostomata) in Galapagos Island fouling communities, Aquatic Invasions 14: 85-131
McCann, Linda; Keith, Inti; Carlton, James T.; Ruiz, Gregory M.; Dawson, Terence P.; Collins, Ken (2015) First record of the non-native bryozoan Amathia (= Zoobotryon) verticillata (delle Chiaje, 1822) (Ctenostomata) in the Galápagos Islands, BioInvasions Records 4: <missing location>
Menon, Ravindranatha; Nair, Balakrishnan (1969) The ectoproctous bryozoans of Indian waters., Journal of the Marine Biological Association of India 9(2): 430-432
Minchin, Dan (2012) Rapid assessment of the bryozoan, Zoobotryon verticillatum (Delle Chiaje, 1822) in marinas, Canary Islands, Marine Pollution Bulletin 64: 2146-2150
Montoya-Cadavid, Erika; Flórez-Romero, Paola; Winston, Judith E. (2007) Checklist of the marine Bryozoa of the Colombian Caribbean, Biota Colombiana 8(2): 159 -184
Mook, David (1983) Indian River fouling organisms, a review, Florida Scientist 26(3/4): 162-167
Nair, N. Balakrishnan (1959) The marine timber-boring molluscs and crustaceans of western Norway, Arbok Universitet i Bergen 25: 1-23
Nair, N. Balakrishnan (1992) Salinity. tolerance in four estuarine species of Bryozoa, Marine Fouling 9(1/2): 15-20
Nascimento, Karine Bianca; Migotto, Alvaro Esteves; Fehlauer Ale, Karin Hoch F (2021) Molecular data suggest the worldwide introduction of the bryozoan Amathia verticillata (Ctenostomata, Vesiculariidae), Marine Biology 168(33): <missing location>
10.1007/s00227-021-03837-8
Oh, Dong-Ha and 6 auhtora (2021) Novel genome characteristics contribute to the invasiveness of Phragmites australis (common reed), Molecular Ecology Resources 31: 1142–1159.
DOI: 10.1111/mec.16293
Orlova, M. I.; Kommendatov, A. Yu. (2013) [The use of laboratory populations of the invasive New Zealand mollusk, Potamopyrgus antipodarum (Gastropoda, Hydrobiidae), for assessment of its euryhalinity and physical modelling of invasion in relation to the salinity gradient], Zoologicheskii Zhurnal 92(7): 759-770
Osburn, R. C. (1914) The Bryozoa of the Tortugas Islands, Florida, Papers from the Tortugas Islands Laboratory of the Carnegie Institution 5: 181-222
Osburn, Raymond C. (1940) Bryozoa of Porto Rico, N. Y. Academy of Sciences - Scientific Survey of Puerto Rico and the Virgin Islands 16(3): 321-486
Osburn, Raymond, C (1953) Bryozoa of the Pacific Coast of America: Part 3, Cyclostomata, Ctenostomata, Entoprocta, and Addenda., Allan Hancock Pacific Expeditions 14(3): 615-684
Otani, Michio (2006) Assessment and Control of Biological Invasion Risks., Shoukadoh Book Sellers and IUCN, Kyoto, Japan, and Gland, Switzerland. Pp. <missing location>
Pederson, E. J.; Peterson, M. S. (2002) Bryozoans as ephemeral estuarine habitat and a larval transport mechanism for mobile benthos and young fishes in the north-central Gulf of Mexico, Marine Ecology Progress Series 140: 935-947
Pestana, L. B.; Dias, G. M.; Marques, A. C . (2021) Spatial and temporal diversity of non-native biofouling species associated with marinas in two Angolan bays, African Journal of Marine Science 42(4): 413-422
Pires-Teixeira, Larissa M. Neres-Lima, Vinicius Creed, Joel C. (2021) How do biological and functional diversity change in invaded tropical marine rocky reef communities?, Diversity 13(353): Published online
Prezant, Robert S.; Toll, Ronald B.; Rollins, Harold B.; Chapman, Eric J. (2002) Marine macroinvertebrate diversity of St. Catherines Island, Georgia., American Museum Novitates 3367: 1-31
Quan-Young, L .I.; Jiménez-Flores, S. G.; Espinoza-Ávalos, J. (2006) [Benthic flora and reproduction of Batophora spp. algae (Chlorophyta: Dasycladaceae) in a polluted coastal lagoon (Chetumal Bay, Mexico)], Revista Biologia Tropical 54(2): 341-355
Quintanilla, Elena; Thomas Wilke; Ramırez-Portilla, Catalina; Sarmiento, Adriana; Sanchez, Juan A. () , None <missing volume>: <missing location>
Rao, K.S.; Ganapati, P. N. (1978) Ecology of fouling bryozoans at Visakhapatnam harbor., Proceedings of the Indian Academy of Sciences <missing volume>: <missing location>
Rizgalla, Jamila; Ashinn, Andrew P.; Crocetta, Fabio (2021) New records of alien and cryptogenic marine bryozoan, mollusc, and tunicate species in Libya, BioInvasions Records 8: 590-597
Ruiz, Gregory; Geller, Jonathan (2021) Spatial and temporal analysis of marine invasions: supplemental studies to evaluate detection through quantitative and molecular methodologies, Marine Invasive Species Program, California Department of Fish and Wildlife, Sacramento CA. Pp. 153 ppl.
Ryland, J. S. (1965) <missing title>, Organization for Economic Co-operation and Development, Paris. Pp. <missing location>
Saenger, P.; Stephenson, W.; Moverley, J. (1979) The subtidal fouling organisms of the Calliope River and Auckland Creek, Central Queensland Australia, Memoirs of the Queensland Museum 19: 399-412
Sambrook, Katie; Holt, Rohan H.F.; Sharp, Rowland; Griffith, Kate; Roche, Ronan C. (2014) Capacity, capability and cross-border challenges associated with marine eradication programmes in Europe: The attempted eradication of an invasive non-native ascidian, Didemnum vexillum in Wales, United Kingdom, Marine Policy 48: 51-59
Schmitt, Joseph D. Emmel, Jason A. Bunch, Aaron J. Hilling, Corbin D. Orth, Donald J (2019) Feeding ecology and distribution of an invasive apex predator: Flathead Catfish in subestuaries of the Chesapeake Bay, Virginia, North American Journal of Fisheries Management 39: 390-402
https://doi.org/10.1002/nafm.1027
Soors, Jan; Faasse, Marco; Stevens, Maarten; Verbessem, Ingrid; De Regge, Nico;Van den Bergh, Ericia (2010) New crustacean invaders in the Schelde estuary (Belgium), Belgian Journal of Zoology 140: 3-10
Soule, Dorothy F.; Soule, John D. (1964) The Ectoprocta (Bryozoa) of Scammon's Lagoon, Baja California, Mexico., American Museum Novitates 2199: 1-56
Soule, Dorothy F.; Soule, John D. (1967) Faunal affinities of some Hawaiian Bryozoa (Ectoprocta), Proceedings of the California Academy of Sciences 35(16): 265-272
Soule, John D. (1963) Results of the Puritan-American Museum of Natural History Expedition to Western Mexico: 18. Cyclostomata, Ctenostomata (Ectoprocta), and Entoprocta of the Gulf of California., American Museum Novitates 2144: 1-34
Tilbrook, Kevin J. (2012) Bryozoa, Cheilostomata: First records of two invasive species in Australia and the northerly range extension for a third, Check List 8(1): 181-183
Tokarev, Yuriy; Shulman, Georgiy (2009) Biodiversity in the Black Sea: effects of climate and anthropogenic factors, Hydrobiologia 580: 23–33
Tovar-Hernández, M. A.; Villalobos-Guerrero, T. F.; Yáñez-Rivera, B., Aguilar-Camacho, J. M.; Ramírez-Santana, I. D. (2012) [Guide to exotic aquatic invertebrates in Sinaloa] , Geomare, A. C., USFWS, INE-SEMARNAT, Mazatlán, México. Pp. 41
U.S. National Museum of Natural History 2002-2021 Invertebrate Zoology Collections Database. http://collections.nmnh.si.edu/search/iz/
Ulman, Aylin and 17 authors (2017) A massive update of non-indigenous species records in Mediterranean marinas, PeerJ 5( e3954): <missing location>
Vail, Lyle L.; Wass, Robin E. (1981) Experimental studies on the settlement and growth of Bryozoa in the natural environment., Australian Journal of Marine and Freshwater Research 32: 639-656
Vaz-Pinto, F. and 5 authors (2014) Invasion success and development of benthic assemblages: Effect of timing, duration of submersion and substrate type, Marine Environmental Research 94: 72-79
Vieira, Leandro M.; Migotto, Alvaro E.; Winston, Judith E. (2008) Synopsis and annotated checklist of Recent marine Bryozoa from Brazil, Zootaxa 1810: 1-39
Waeschenbach, Andrea; Vieira, Leandro M.; Reverter-Gil, Oscar; Souto-Derungs, Javier; Nascimento, Karine B.; Fehlauer-Ale, Karin H. (2015) A phylogeny of Vesiculariidae (Bryozoa, Ctenostomata) supports synonymization of three genera and reveals possible cryptic diversity, Zoologica Scripta <missing volume>: <missing location>
Webb, Alan Charles (2007) Status of non-native freshwater fishes in tropical northern Queensland, including establishment success, rates of spread, range and introduction pathways, Journal and Proceedings of the Royal Society of New South Wales 149: 63-78
Weiss, Charles M. (1948) The seasonal occurrence of sedentary marine organisms in Biscayne Bay, Florida., Ecology 29(2): 153-172
Williams, Susan L. (2007) Introduced species in seagrass ecosystems: Status and concerns., Journal of Experimental Marine Biology and Ecology 350: 89-110
Winston, Judith E. (1977) Distribution and ecology of estuarine ectoprocts: a critical review., Chesapeake Science 18(1): 34-57
Winston, Judith E. (1982) Marine bryozoans (Ectoprocta) of the Indian River area (Florida)., Bulletin of the American Museum of Natural History 173: 99-176
Winston, Judith E. (1984) Shallow-Water Bryozoans of Carrie Bow Cay, Belize, American Museum Novitates 2799: 1-38
Winston, Judith E. (2004) Bryozoans from Belize, Atoll Research Bulletin 523: 1-14
Winston, Judith E. (2010) Stability and change in the Indian River area bryozoan fauna over a twenty-four year period, Smithsonian Contributions to the Marine Sciences 38: 229-239
Wirtz, Peter 2/3/2021 <em>Amathia verticillata</em> (Bryozoa, Ctenostomata) at the coast of Senegal.. <missing URL>
Wirtz, Peter; Canning-Clode, João (2009) The invasive bryozoan Zoobotryon verticillatum has arrived at Madeira Island, Aquatic Invasions 4(4): 669-670
Woods Hole Oceanographic Institution, United States Navy Dept. Bureau of Ships (1952) Marine fouling and its prevention., United States Naval Institute., Washington, D.C.. Pp. 165-206
Wyatt, Alex S. J.; Hewitt, Chad L.; Walker, Di I.; Ward, Trevor J. (2005) Marine introductions in the Shark Bay world heritage property, Western Australia: a preliminary assessment., Diversity and Distributions 11: 33-44
Xavier, Everthon A; Aalmeida, na C.S.; Vieira, Leandro M. (2021) The role of artificial habitats on fouling bryozoan fauna in the southwestern Atlantic, Marine Pollution Bulletin 167: Published online
https://doi.org/10.1016/j.marpolbul.2021.112310
Zambrano, René; Ramos, John (2021) Alien crustacean species recorded in Ecuador, Nauplius 29: e2021043