Invasion History

First Non-native North American Tidal Record: 1977
First Non-native West Coast Tidal Record: 1977
First Non-native East/Gulf Coast Tidal Record: 1999

General Invasion History:

Ianiropsis serricaudis was described from the Pacific coast of Russia (Gurjanova 1936, cited by Hobbs et al. 2015), and is known to occur from the Korea Straits to the Sea of Okhotsk (Jang and Kwon 1990; Hobbs et al. 2015). In 1977, it was tentatively identified from San Francisco Bay by Carlton, Iverson, and Chapman (Carlton 1979), but was later not considered to be established because of taxonomic uncertainty (Cohen and Carlton 1995). In 2000, an unidentified Ianiropsis, the first of the genus to be found on the East Coast, was found in rapid assessment surveys of nonindigenous species in New England (MIT Sea Grant 2000-2008; Pederson et al. 2003). Collections of similar isopods were made from fouling communities in the Netherlands (in 2000, Faasse 2007, cited by Hobbs et al. 2015); England (in 2004, Hobbs et al. 2015); Puget Sound (in 2010, Cordell et al. 2012), and the Lagoon of Venice (in 2012, Marchini et al. 2015). Likely vectors for the spread of this isopod are ship fouling, ballast water, and oyster aquaculture (Hobbs et al. 2015).

North American Invasion History:

Invasion History on the West Coast:

Ianiropsis serricaudis was collected and identified from fouling communities in San Francisco Bay by Carlton, Iverson, and Chapman in 1977. It was found associated with the introduced tunicates Styela clava, Ciona robusta, and the isopod Dynoides dentisinus (Carlton 1979). Ianiropsis spp. continued to be collected, but identifications to species were difficult, owing to a large number of native species. Cohen and Carlton (1995) listed I. serricaudis among species whose establishment was uncertain. Specimens collected in the 1990s and 2000s from Treasure Island (1993), Richmond (2004), and the Presidio Yacht Club (2004) were verified as I. serricaudis (Hobbs et al. 2015), all in the central Bay. Additional collections were made in 2005 at Coyote Point Marina and San Mateo, in South San Francisco Bay (Foss 2009). In 2002, three specimens of I. serricaudis were identified from Elkhorn Slough, in Moss Landing. The isopods were found near reefs of the tubeworm Ficopomatus enigmaticus (Heiman and Micheli 2010; Hobbs et al. 2015). This isopod was reported from open coast areas at Point Arena and Purisima Point in 2004 (Maloney et al. 2006), but we do not know if populations are established there.

In 2010, I. serricaudis was found to be abundant in Puget Sound, Washington at one site: Taylor Shellfish Farms in Totten Inlet (Cordell et al. 2012). It was associated with Didemnum vexillum there, but was not found at other locations in Puget Sound (Cordell et al. 2012; Hobbs et al. 2015). Living specimens of I. serricaudis were found on a dock and a boat that drifted across the Pacific to the Oregon coast after the Japanese earthquake and tsunami of 2011, but these transports are not known to have resulted in established populations (Hobbs et al. 2015).

Invasion History on the East Coast:

There are no native Ianiropsis spp. in the Northwest Atlantic, but the small size of I. serricaudis makes it easy to overlook. On the East Coast, Ianiropsis serricaudis was first collected in 1999 in the Mystic River, a tributary of Long Island Sound, 'amongst the native bryozoan Amathia dichotoma', but not identified until 2012 (Hobbs et al. 2015). In New England, surveys conducted in 2000-2014 found I. serricaudis, identified as Ianiropsis sp., from Waterford, Connecticut to Bourne Marina, Massachusetts (MA) (MIT Sea Grant 2003-2008; Pederson et al. 2003; Janiak and Whitlach 2012). In 2007-2013, it was found in the Gulf of Maine, from Sandwich, MA (Cape Cod Bay) to Camden, Maine, (Penobscot Bay) (MIT Sea Grant 2009-2012; Wells et al. 2014). It was typically found on dock floats overgrown with Ulva sp. and tunicates, and rocky intertidal shores (Janiak and Whitlach 2012; Hobbs et al. 2015). In 2012-2013, I. serricaudis was found in Barnegat Bay, New Jersey, associated with the tunicate Botryllus schlosseri (Pallas, 1766), and the native bryozoans Einhornia crustulenta (Pallas, 1766) and Schizoporella sp., as well as the green alga Ulva lactuca (Hobbs et al. 2015). In 2012, this isopod was found in at one station in Sinepuxent Bay, Maryland, at only one of 13 stations in the Maryalnd Coastal Bays (Morales-Núñez, and Chigbu 2018).

Invasion History Elsewhere in the World:

In Europe, I. serricaudis was first observed in the Oosterschelde, the Netherlands in 2000 (Faasse 2007, cited by Hobbs et al. 2015). It was later found at another site, near the port of Rotterdam. In England, it was found in 2004 at Southampton, in the Solent estuary, off the English Channel (Hobbs et al. 2015), and at Plymouth, on the southwest coast (Hobbs et al. 2015). In 2012, it was found in the Lagoon of Venice, its first reported occurrence in the Mediterranean (Marchini et al. 2015). As with the invasions in North America, hull fouling and ballast water are probably the likeliest modes of long-distance transport, but aquaculture sites are likely areas of colonization and a source of local dispersal (Hobbs et al. 2015).


Description

Ianiropsis serricaudis has a lozenge-shaped body, about 3X as long as wide, with the sides nearly parallel, broadest in pereaonal segment 3, and slightly tapered towards the head. The margin of the cephalon is slightly concave and has an inconspicuous rostrum on the frontal margin. The pleotelson has a small median lobe, with 3 or 4 spine-like serrations on each side (sometimes up to 7) and 2 or 3 unequal setae at the base of each serration. The uropods are slightly longer than the pleotelson. Antenna 1 is comparatively short, reaching segment 5 of Antenna 2 and about 1/4 of the body length. The Antennae 2 of I. serricaudis are about equal to body length, and peduncle segments 6 and 7 are especially elongated, totaling about 1/2 body length. In mature males, the maxilliped palps are elongated, with 5 segments, extending well beyond the basal segments of Antenna 2 and visible in dorsal view. The pereiopods are long, 1X-2X body maximum width. Pereiopods 1 have 3 claws each, while Pereiopods 7 have 2-3 claws. Antenna 2 is 6/10 of body length, with a flagellum of 24 segments. In males, Pleopod pairs 1 and 2 are modified for copulation. Pleopods 1 are fused to form a narrow appendage (the sympod),widening near the tip and ending in a pair of thick spines, each bearing 4 outer setae, a group of short setae on the inner side of the spine, and a longer, medial group of 6-7 longer setae. Pleopods 2 are not fused, but the inner ramus of each is modified into a needle-like stylus. Males are up 3.2 mm long and females are at least 2.4 mm. Reported colors vary, apparently with substrate. Gurjanova (1936, cited by Hobbs et al. 2015), reported that the isopods in Pacific Russia were pale, but with black eyes, spotted with black melanophores, giving a gray appearance. Korean specimens had brown melanophores (Jang and Kwon 1990), while Dutch animals often had red eyes and brown melanophores. New England animals varied from greenish to yellowish, and reddish, according to the color of the bryozoans and sponges on which they were feeding. Description based on: Jang and Kwon 1990, Wilson and Wagele 1994, Doti and Wilson 2010, Hobbs et al. 2015.


Taxonomy

Taxonomic Tree

Kingdom:   Animalia
Phylum:   Arthropoda
Subphylum:   Crustacea
Class:   Malacostraca
Subclass:   Eumalacostraca
Superorder:   Peracarida
Order:   Isopoda
Suborder:   Asellota
Family:   Janiridae
Genus:   Ianiropsis
Species:   serricaudis

Synonyms

Ianiropsis sp. (Pederson et al., 2003)
Ianiropsis notoensis (Nunomura, 1985)

Potentially Misidentified Species

Ianiropsis analoga
Menzies 1952, Northeast Pacific (Schultz 1969; Brusca et al. 2007; Hobbs et al. 2015)

Ianiropsis breviremis
Sars 1883, Northeast Atlantic (Hobbs et al. 2015)

Ianiropsis derjugini
Gurjanova 1952, Northwest Pacific, Bering Sea, Northeast Pacific (Jang and Kwon 1990; Brusca et al. 2007; Hobbs et al. 2015)

Ianiropsis epilittoralis
Menzies 1952, Northwest Pacific, Northeast Pacific (Schultz 1969; Jang and Kwon 1990; Brusca 2007; Hobbs 2015)

Ianiropsis kincaidi
Richardson 1904, Northeast Pacific (Schultz 1969; Brusca 2007)

Ianiropsis minuta
Menzies 1952, Northeast Pacific (Schultz 1969; Brusca 2007)

Ianiropsis montereyensis
Menzies 1952, Northeast Pacific (Schultz 1969; Brusca 2007)

Ianiropsis tridens
Menzies 1952, Northwest Pacific, Northeast Pacific, Chile (Schultz 1969; Jang and Kwon 1990; Brusca 2007; Hobbs et al. 2015)

Ecology

General:

Ianiropsis serricaudis has separate sexes, with internal fertilization, and direct development. Kussakin (1988, cited by Hobbs et al. 2015) observed breeding of I. serricaudis in Russia, in August-October, with 7-32 (average 18) eggs per female.

Ianiropsis serricaudis occurs in cold-temperate to warm-temperate climates, and has been collected at temperatures ranging from -1.8 C to 24 C. It has been collected at salinities from 16.9 to 35 PSU (Hobbs et al. 2015). This isopod occurs on a wide range of fouling organisms which provide shelter, including algae (red algae-Pachyarthron cretaceum, Chondrus crispus, Grateloupia turuturu; brown algae- Laminaria sp., green algae- Ulva spp.); sponges (Halichondria bowerbanki, Clathria prolifera); bryozoans (Amathia dichotoma, Einhornia crustulenta, Schizoporella sp., Tricellaria inopinata); and tunicates (Botrylloides violaceus, Ciona robusta, Didemnum vexillum, Diplosoma listerianum, Styela clava). Many of the organisms used by I. serricaudis are native to the Northwest Pacific, the native region of I. serricaudis (Hobbs et al. 2015). The feeding habits of this isopod is not known, but it may feed on the fecal material of colonial invertebrates, as well as, the surface microbiota and detritus associated with colonial invertebrates and algae.

Food:

Microalgae, protozoans, pseudofeces

Trophic Status:

Omnivore

Omni

Habitats

General HabitatMarinas & DocksNone
General HabitatRockyNone
Salinity RangePolyhaline18-30 PSU
Salinity RangeEuhaline30-40 PSU
Tidal RangeSubtidalNone
Vertical HabitatEpibenthicNone


Tolerances and Life History Parameters

Minimum Temperature (ºC)-1.8Field, Pacific Russia (Hobbs et al. 2015)
Maximum Temperature (ºC)24Field, Pacific Russia, Narragansett Bay (Hobbs et al. 2015)
Minimum Salinity (‰)16.9Field (Hobbs et al. 2015)
Maximum Salinity (‰)35Field (Hobbs et al. 2015)
Maximum Length (mm)3.2Hobbs et al. 2015
Broad Temperature RangeNoneCold-temperate
Broad Salinity RangeNonePolyhaline-Euhaline

General Impacts

Impacts of Ianiropsis serricaudis have not been studied. However, in some cases this isopod can reach extraordinary abundances within fouling communities, and could have impacts as a competitor, as prey, or in some other ecological role (Hobbs et al. 2015).

Regional Distribution Map

Bioregion Region Name Year Invasion Status Population Status
NEP-IV Puget Sound to Northern California 2015 Non-native Established
NEP-V Northern California to Mid Channel Islands 2013 Non-native Established
P116 _CDA_P116 (Big Navaro-Garcia) 2004 Non-native Unknown
P067 _CDA_P067 (San Antonio) 2004 Non-native Unknown
P080 Monterey Bay 2002 Non-native Established
P090 San Francisco Bay 1977 Non-native Established
NEP-V Northern California to Mid Channel Islands 1977 Non-native Established

Occurrence Map

OCC_ID Author Year Date Locality Status Latitude Longitude
756175 Introduced Species Study 2005 2005-06-08 Sea Plane Lagoon Non-native 37.7761 -122.2998
756176 Introduced Species Study 2005 2005-06-09 Point Cavallo Non-native 37.8319 -122.4737
756177 Introduced Species Study 2005 2005-08-25 Ferry Terminal Pier Non-native 37.7945 -122.3917
756178 Introduced Species Study 2005 2005-08-25 China Basin Non-native 37.7780 -122.3881
756179 Introduced Species Study 2005 2005-08-25 Potrero Point Non-native 37.7521 -122.3790
756180 Introduced Species Study 2005 2005-09-07 Dumbarton Bridge Non-native 37.5070 -122.1168
756181 Introduced Species Study 2005 2005-09-07 Redwood Creek - Marina Non-native 37.5021 -122.2130
756182 Introduced Species Study 2005 2005-09-07 Railroad Bridge Non-native 37.4602 -121.9750
756183 Introduced Species Study 2005 2005-09-08 Treasure Island Non-native 37.8149 -122.3702
756184 Introduced Species Study 2005 2005-09-08 Pier 39 Non-native 37.8108 -122.4086
756185 Introduced Species Study 2005 2005-09-08 Pier 45 Non-native 37.8111 -122.4196
756186 Introduced Species Study 2005 2005-09-08 Cruise Ship Pier Non-native 37.8085 -122.4060
756187 Introduced Species Study 2005 2005-09-09 Sea Plane Harbor Non-native 37.6349 -122.3848
756188 Introduced Species Study 2005 2005-09-09 San Mateo Bridge Non-native 37.5806 -122.2543
756189 Introduced Species Study 2005 2005-09-09 Sierra Point Marina Non-native 37.6740 -122.3792
756190 Introduced Species Study 2005 2005-09-09 Coyote Point Marina Non-native 37.5905 -122.3177
756191 Introduced Species Study 2005 2005-10-04 Oakland Outer Harbor Non-native 37.8217 -122.3145
756192 Introduced Species Study 2005 2005-10-04 Oakland Inner Harbor - Shipping cranes Non-native 37.7947 -122.3095
756193 Introduced Species Study 2005 2005-10-04 Berkeley Flats/Berkeley Pier Non-native 37.8600 -122.3256
756194 Introduced Species Study 2005 2005-10-04 Berkeley Marina Non-native 37.8676 -122.3172
756195 Introduced Species Study 2005 2005-10-05 Oakland Inner Harbor - Small marinas Non-native 37.7847 -122.2669
756196 Introduced Species Study 2005 2005-10-05 Port of Oakland Office Non-native 37.7954 -122.2804
756197 Introduced Species Study 2005 2005-10-05 Sea Plane Lagoon Non-native 37.7761 -122.2998
756198 Introduced Species Study 2005 2005-10-05 Coast Guard Island Non-native 37.7812 -122.2457
756199 Introduced Species Study 2005 2005-10-05 Ballena Bay Non-native 37.7661 -122.2834
756200 Introduced Species Study 2005 2005-10-06 Point Richmond Piers Non-native 37.9085 -122.3913
756201 Introduced Species Study 2005 2005-10-06 Richmond Marina Non-native 37.9137 -122.3504
756202 Introduced Species Study 2005 2005-10-06 Santa Fe Channel - Back Non-native 37.9207 -122.3684
756203 Introduced Species Study 2005 2005-10-06 Santa Fe Channel - Front Non-native 37.9101 -122.3644
756204 Introduced Species Study 2005 2005-10-19 Mare Island Strait - Navy Non-native 38.1015 -122.2695
756205 Introduced Species Study 2005 2005-10-19 Hercules Wharf Non-native 38.0231 -122.2928
756206 Introduced Species Study 2005 2005-10-19 Mare Island Strait - Marina Non-native 38.1051 -122.2667
756207 Introduced Species Study 2005 2005-10-20 Point San Pablo Yacht Harbor Non-native 37.9643 -122.4185
756208 Introduced Species Study 2005 2005-10-20 Port Sonoma/Petaluma R. Non-native 38.1157 -122.5026
756209 Introduced Species Study 2005 2005-10-20 San Pablo Bay Pumphouse Non-native 38.0446 -122.4326
756210 Introduced Species Study 2005 2005-10-21 Ayala Cove Non-native 37.8680 -122.4350
756211 Introduced Species Study 2005 2005-10-21 Corinthian Marina Non-native 37.8726 -122.4563
756212 Introduced Species Study 2005 2005-10-21 Richardson Bay Non-native 37.8588 -122.4798
756213 Introduced Species Study 2010 2010-05-31 Railroad Bridge Non-native 37.4602 -121.9750
756214 Introduced Species Study 2010 2010-05-31 Dumbarton Bridge Non-native 37.5070 -122.1168
756215 Introduced Species Study 2010 2010-06-02 Ballena Bay Non-native 37.7661 -122.2834
756216 Introduced Species Study 2010 2010-06-02 Oakland Inner Harbor - Shipping cranes Non-native 37.7947 -122.3095
756217 Introduced Species Study 2010 2010-06-02 Port of Oakland Office Non-native 37.7954 -122.2804
756218 Introduced Species Study 2010 2010-06-03 Treasure Island Non-native 37.8149 -122.3702
756219 Introduced Species Study 2010 2010-06-03 Berkeley Marina Non-native 37.8676 -122.3172
756220 Introduced Species Study 2010 2010-06-28 Santa Fe Channel - Back Non-native 37.9207 -122.3684
756221 Introduced Species Study 2010 2010-06-30 Hercules Wharf Non-native 38.0231 -122.2928
756222 Introduced Species Study 2010 2010-06-30 Mare Island Strait - Marina Non-native 38.1051 -122.2667
756223 Introduced Species Study 2010 2010-07-01 Richardson Bay Non-native 37.8588 -122.4798
756224 Introduced Species Study 2010 2010-07-01 Ayala Cove Non-native 37.8680 -122.4350
756225 Introduced Species Study 2010 2010-07-12 Pier 45 Non-native 37.8111 -122.4196
756226 Introduced Species Study 2010 2010-07-13 Port Sonoma/Petaluma R. Non-native 38.1157 -122.5026
760480 Carlton 1979a; Hobbs et al. 2015 1977 Oakland Estuary Non-native 37.7866 -122.2654
760481 Hobbs et al. 2105 1993 Treasure Island Yacht Harbor, Clipper Cove Non-native 37.8158 -122.3701
760482 Hobbs et al. 2105 2004 Richmond Marina Non-native 37.9137 -122.3504
760483 Hobbs et al. 2105 2004 Presidio Yacht Club (Horseshoe Bay) Non-native 37.8327 -122.4741
760484 Heiman and Micheli 2010; Hobbs et al. 2015 2002 Elkhorn Slough (Upper) Non-native 36.8547 -121.7600
767353 Ruiz et al., 2015 2012 2012-08-21 Porto Bodega, Bodega Bay, California, USA Non-native 38.3333 -123.0525
767375 Ruiz et al., 2015 2012 2012-08-21 Tomales-Nick's Cove, Bodega Bay, California, USA Non-native 38.1980 -122.9222
767394 Ruiz et al., 2015 2012 2012-08-16 Tomales-SNPS, Bodega Bay, California, USA Non-native 38.1359 -122.8719
767604 Ruiz et al., 2015 2013 2013-09-05 Launch Ramp, Morro Bay, CA, California, USA Non-native 35.3577 -120.8508
768191 Ruiz et al., 2015 2012 2012-09-07 Jack London Square Marina, San Francisco Bay, CA, California, USA Non-native 37.7940 -122.2787
769437 Ruiz et al., 2021b 2018 2018-08-10 Alamitos Bay Marina, San Pedro Bay, California, USA Non-native 33.7494 -118.1154
770034 Ruiz et al., 2021b 2018 2018-08-08 Port of LB Pier F, San Pedro Bay, California, USA Non-native 33.7472 -118.2130

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